Caprifoliaceae Juss.

Honeysuckle family (en), Caprifoliacées (fr)

Family

Angiosperms > Dipsacales

Characteristics

Monoecious, dioecious (not in Australia), polygamous (not in Australia) or gynodioecious (not in Australia) annual, biennial or perennial herbs, shrubs or lianas. Leaves opposite, sometimes whorled or in a basal rosette, simple or lobed, entire to pinnately dissected, petiolate; stipules absent, sometimes with pseudostipules. Inflorescences terminal (not in Australia) or axillary, thyrses, spikes, racemes or heads. Flowers bisexual or female, actinomorphic or zygomorphic, sometimes with an epicalyx of 4 fused and modified bracts. Sepals fused, forming a cup or tube, with 4 or 5 lobes, sometimes modified into spiny setae or further divided into up to 30 pappus-like segments. Petals fused, 4 or 5, infundibular, rotate or campanulate, usually bilabiate. Nectary usually 1 in corolla tube at base of abaxial petal or rarely 3–5. Stamens 1–5, epipetalous, alternating with corolla lobes; anthers 2-locular, dehiscing by longitudinal slits. Ovary inferior, 2–5-or rarely 7–10-carpellate, 1–3-locular; ovules one to numerous per locule; placentation axile. Fruit a many seeded septicidal capsule (not in Australia) or berry, drupe with 2–4 pyrenes or an achene or nutlet enclosed in an epicalyx or often with a persistent, sometimes pappus-like or awned calyx; seeds with or without endosperm.
More
Shrubs or woody climbers, rarely small trees or herbs. Leaves opposite, rarely whorled, simple or pinnatifid, conduplicate or involute in vernation; interpetiolar stipules absent or rarely well developed. Inflorescence thyrsoid, axillary or terminal, compact or lax, cymes 1-, 2-, or 3-flowered; paired flowers sometimes with ovaries ± fused. Cymes with a pair of bracts and 2 pairs of bracteoles, located at base of ovaries, ± fused, occasionally accrescent in fruit, rarely absent. Flowers bisexual, actinomorphic or zygomorphic. Calyx 4-or 5-lobed. Corolla epigynous, gamopetalous; lobes 4 or 5, spreading, sometimes bilabiate, aestivation imbricate. Stamens (4 or)5, didynamous, alternating with corolla lobes, sometimes exserted; anthers free, 2-celled, opening by longitudinal slits, introrse. Ovary inferior, carpels 2-8, fused; placenta axile; ovules 1 to many per locule, some of which can be abortive, pendulous; style solitary; stigmas capitate or lobed. Fruit a berry, a drupe with 2-5 pyrenes, or a leathery achene. Seeds 1 to many; embryo small, straight; endosperm copious.
Small trees, shrubs, woody climbers or rarely herbs, often with soft pith. Leaves opposite or less often alternate, simple or pinnately compound; stipules absent or very small, rarely conspicuous. Inflorescences mainly cymose, less often flowers solitary or paired; bracts and bracteoles usually present. Flowers regular or irregular, hermaphrodite. Calyx-tube joined with the ovary; limb (3–)4–5-fid or-toothed, the teeth imbricate or open. Corolla gamopetalous, tubular, funnel-shaped, campanulate or rotate, 3–5-lobed, sometimes 2-lipped, the lobes imbricate or less often valvate. Stamens 4–5, inserted in the corolla-tube and alternate with the lobes; anthers 2-thecous, opening lengthwise inwards or outwards. Ovary inferior, 2–8-locular, with 1-many pendulous ovules in each locule; placentation axile or parietal; style simple or wanting; stigma ], capitate, or stigmas as many as the carpels. Fruit a berry or drupe, less often a capsule or achene, 1–5-locular with 1–many seeds or as many pyrenes as locules; endosperm copious; embryo mostly small and straight
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Hardiness (USDA) 8-11

Usage

Several of the genera, including Abelia, Centranthus, Lonicera and Symphoricarpos are widely cultivated. The traditional Chinese medicine Xu Duan is made from the roots of Dipsacus and is used to combat joint and bone ailments as well as external sores and uterus bleeding (Mayer 2016). In South Africa the leaves or roots of Scabiosa columbaria are used to remedy colic and heartburn and to heal wounds (Mayer 2016). Rhizomes of Nardostachys jatamansi and Valeriana celtica are used to extract oils to serve as stimulants (Weberling & Bittrich 2016). The leaves of Valerianella species and Valeriana edulis are consumed in Europe and Mexico, respectively (Weberling & Bittrich 2016).
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