Rhizome c. 1.5-2.6 mm thick, long-creeping, phyllopods 2-7 cm apart, buds halfway along the internodes. Anatomy: vascular strands 5-11, sclerenchyma sheath distinct, sclerenchyma strands c. 15, in a ring within the vascular cylinder, often confluent. Rhizome scales spreading to recurved, peltate, 5-11 by 0.5-1 mm, base entire, acumen entire to dentate, hyaline. Fronds dimorphic, stalked. Fertile fronds with stalks 0.5-15(-25) cm long, to 0.5(-0.75) F as long as the lamina, lamina 8-44 by 1-3.5 cm, index 5-20 or more, base narrowed, widest below middle, apex acute to acuminate. Sterile fronds with stalks 2-9 cm, lamina 5-22 by 1.5-4.5 cm, index 2-7, widest near the middle, otherwise similar to the fertile fronds. Venation: veins distinct, veinlets branched and often anastomosing, free veinlets mostly excurrent. Hydathodes absent. Stellate hairs dimorphic, forming a thin to thick mat, often fugaceous, light whitish brown, the straight upper rays boat-shaped to acicular. Sori apical to all over the lamina, 3-4 mm wide, often elongate, slightly sunken, developing acropetally. Sporangia long-stalked, paraphyses with short, straight rays. Spores with longitudinal ridges.