Rhizome (1-)1.5-2.6 mm thick, long-creeping, phyllopods 2-7 cm apart, buds halfway along the internodes. Anatomy: vascular strands 6-10, sclerenchyma sheath distinct, sclerenchyma strands c. 10, one central, the other peripheral. Rhizome scales spreading, peltate, 3.2-7.2 by 0.6-1.5 mm, base entire to dentate, acumen long-ciliate in the lower half, light brown, apex entire, crisped. Fronds dimorphic, stalked. Fertile fronds with stalks to 3-16 cm long, 0.3-1 as long as the lamina, lamina 5-16 by 1-3 cm, index (2-)4-8, base cuneate to gradualy attenuate, widest at or below the middle, apex obtuse to acuminate. Sterile fronds with stalks 1-14 cm, 0.1-1 as long as the lamina, lamina 3.5-17 by 1.3-6 cm, index 2-4(-5), base cuneate to attenuate, apex obtuse or more or less rounded to long-acuminate. Venation: veins distinct, veinlets much branched and copiously anastomosing, free veinlets many, excurrent and recurrent. Hydathodes distinct. Stellate hairs monomorphic, appressed, forming a thin mat, light brown to whitish, the rays short, boat-shaped. Sori all over the lamina or in an irregular, sharply defined patch, very closely packed, superficial, developing more or less simultaneously, soral area appearing acrostichoid when old. Sporangia stalked, paraphyses not differentiated. Spores with irregular ridges and protuberances.