Trees or shrubs, unarmed. Leaves simple, petiolate, the blade generally entire, sometimes cordate at the base, penninerved or palminerved. Flowers axillary or op-positifolious, sometimes cauliflorous, solitary or fasciculate, hermaphrodite, actino-morphous, pedicellate, generally (2-)3-bracteolate, the bracteoles caducous or per- sistent; receptacle eglandular; calyx campanulate or tubiform, irregularly dentate to lobate, sometimes longitudinally winged, coriaceous, lepidote or stellate-pubescent outside, accrescent; petals 5, adnate to the base of the staminal column, fleshy; sta-mens numerous, the staminal column elongate, undivided to 5-dentate or 5-lobate, the anthers sessile or subsessile near the apex of the staminal column or on the lobes, 1-thecate, the thecae paired or not, free or sometimes more or less confluent at the apex, extrorse, longitudinally dehiscent; pollen porate or colporate, the sexine foveolate or ornate-reticulate; ovary subinferior, 2-or 4-or 5-celled, the cells 2-ovulate; style simple; stigma disk-shaped, sometimes shallowly sulcate or lobulate. Capsules drupaceous, indehiscent, the mesocarp coriaceous or fleshy-pulpy, 1-to 5-celled; seeds generally 1. in each cell; cotyledons fleshy and contortuplicate.