Plants medium-sized to large, coarse and rigid, usually hoary, grayish, brownish or yellowish green, yellow or yellow-to blackish brown. Stems mostly pinnately branched, with many short, lateral branchlets. Leaves erect to slightly secund when dry, loosely erect to erect-spreading or spreading-recurved when wet, narrowly ovate-to linear-lanceolate; margins 1-stratose, recurved to revolute, entire proximally, coarsely dentate along the hyaline border; apices gradually tapering to a long, slender, hyaline acumen; awns densely papillose, erose-dentate, long-decurrent, with the decurrencies flat or ruffled; costa percurrent, in transverse section 2-stratose, becoming 3-stratose in the proximal portion; laminal cells 1-stratose, sinuose, dull and opaque, distinctly papillose with large, flat papillae covering the longitudinal walls and almost the whole of the lumina except for a narrow central groove; basal marginal laminal cells long-rectangular, forming 1(-2)-seriate band, consisting of to 30 rectangular, translucent, not sinuose cells; alar cells not differentiated; medial cells long-rectangular; distal cells short-rectangular. Inner perichaetial leaves hyaline, oblong, oblong-lanceolate to elliptical, abruptly constricted into a short, smooth or weakly papillose awn. Seta single or often 2-3 per perichaetium, sinistrorse when dry, strongly papillose. Capsule straight, ovoid to ovoid-cylindric, somewhat ventricose in the base; annulus revoluble, 2-4-seriate; operculum long-rostrate; peristome teeth, long, reddish brown, split nearly to the base into 2 filiform, strongly papillose branches. Calyptra conic-mitrate to cucullate, naked. Spores spherical, pale yellow, finely roughened.
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