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Female flowers fewer than male and coaxillary with them; perianth similar to that of male flower, the receptacle-lobes usually smaller and less divided; ovary densely setose; ovules many, horizontal; stigma 3-lobed.
Fruit subglobose to ellipsoid-cylindrical, fleshy, densely echinate with multicellular brownish setae, dehiscent by 10 apical valves and extruding seeds and placentae.
Receptacle-tube short, obconic-campanulate; lobes lanceolate to broadly foliaceous, often pinnately divided into narrow or ± expanded often deflexed lobes.
Stamens 3, all two-thecous; filaments inserted on upper part of receptacle-tube; thecae flexuous.
Seeds asymmetrically broadly ovate in outline, compressed; testa smooth, bordered.
Male flowers in sessile axillary fascicles, bracteate.
Corolla gamopetalous, 5-lobed, lobes entire.
Flowers yellow, monoecious.
Leaves simple, petiolate.
Probracts present.
Tendrils simple.
Scandent herbs.