Annual, biennial, or perennial herbs; pubescence simple; stems single from the base, often coarse, usually branched above, glabrous or pubescent; basal leaves present or absent; cauline leaves sessile, auriculate, sagittate, usually longer than the internodes; inflorescences greatly elongated, terminating many branches as well as the main stem; flowers mostly numerous, small; sepals caducous or at least early deciduous, non-saccate; petals narrow, spreading at anthesis; stamens equal or nearly so, mostly spreading at anthesis, filaments enlarged at base; nectar-glands mostly well developed, surrounding or at least subtending all stamens; siliques terete to slightly flattened parallel to replum, erect, widely spreading or deflexed, stipitate to nearly sessile; seeds wingless, plump to somewhat flattened, uniseriate to biseriate; cotyledons incumbent to accumbent.