Tree, monoecious. Indumentum of solitary, simple, densely appressed, short brown hairs only. Twigs glabrous except for the tip. Leaves paripinnate, unifoliolate or 1-or 2-jugate (Malesian species); no pseudo-stipules; neither petiole nor rachis winged; petiolules strongly swollen. Leaflets opposite (Malesian species), not papillose, (sub)glabrous, without domatia or red glands; base symmetrical, acute; margin entire (except for 2 Aus-traliar species); midrib above prominulous, beneath prominent; nerves widely spreading. Inflorescences axillary and sometimes ramiflorous, racemose, simple or with 1 or 2 basal branches. Flowers unisexual, actinomorphic. Sepals 5, free, slightly imbricate, equal, not petaloid, outside sparsely, inside densely hairy. Petals 5 or 0, shorter than the sepals, outside near the base sparsely hairy, inside long-hairy in the lower half, with woolly auricles, no crest. Disc complete, densely hairy (New Guinea) or glabrous (Australia). Stamens (5-)7 (or 8), hardly exserted; filaments hairy; anthers glabrous. Pistil hairy; ovary 2-or 3-locular with 1 ovule per locule; smooth, not lobed; style apical, about as long as the ovary; stigma slightly lobed. Fruits capsular, dehiscence loculicidal, completely 2-or 3-celled, slightly 2-or 3-lobed, not winged, shortly stipitate, outside ± smooth, very sparsely shortly sericeous, pericarp ± fleshy, completely 2-or 3-celled, inside densely woolly. Seeds ellipsoid, glossy blackish brown; sarcotesta annular around the hilum to cupukr and covering up to about a third part of the seed.
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Monoecious trees. Indumentum of simple hairs. Leaves paripinnate; leaflets opposite or alternate, usually decreasing in size from apex to base. Inflorescence axillary or ramiflorous, usually slender, raceme-like or openly branched. Flowers regular. Calyx cupular, with 5 ovate lobes. Petals 5, usually ovate, clawed, the lower margins incurved or auricled; scales 2, hairy, crestless. Disc annular. Stamens 8; filaments filiform, hairy in lower half. Ovary 2-or 3-locular; ovule 1 per locule, 1 or 2 mostly abortive; style slender. Fruit ellipsoidal, obovoid or subglobose, 1–3-locular, loculicidally dehiscent, 2-or 3-valved; pericarp usually fleshy but valves drying thin, usually hairy inside. Seed usually with thin, shiny, brittle testa; aril basal, shortly cupular or rudimentary, rarely absent.
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