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Annual or perennial glandular-pilose herbs; stems erect. Leaves opposite, sessile or with winged petiole simple, entire, toothed or lobed. Capitula small, in terminal cymes, broadly campanulate or hemispherical; involucral bracts 1-seriate, herbaceous, 5, spreading or reflexed; receptacle convex to short-conical; outer paleae cucullate, green to purplish, each enclosing a ray floret; inner paleae membranous, becoming flatter, white to purplish. Ray florets in 1 series, female, fertile, 3-toothed, inconspicuous. Disc florets bisexual, fertile, tubular, 5-toothed. Achenes obovoid-oblong, often incurved. Pappus absent.
Annual herbs. Lvs opposite, simple, entire to serrate or incised. Capitula pedunculate, solitary or in loose panicles. Involucral bracts in 2 rows; outer row foliaceous, glandular; inner row smaller, partly membranous. Receptacle flat; scales present. Outer florets ♀, shortly ligulate, yellow. Inner florets ☿, tubular. Achenes all similar, square or triangular in section, narrowed to base, truncate at apex; pappus 0.
Annual or perennial herbs. Leaves opposite. Capitula in lax panicles, heterogamous; involucre of 3–5 phyllaries with stalked glands; receptacle paleate. Ray florets 1-seriate, female, inconspicuous; disc florets hermaphrodite, 3–5-lobed; anthers with minute apical appendage; style branches short and flattened. Achenes usually incurved; pappus absent.
