Herbs annual, biennial, or perennial, or subshrubs. Trichomes dendritic, sometimes mixed with fewer simple and stalked forked ones; glandular trichomes multicellular, multiseriate. Stems erect, simple or branched. Basal leaves petiolate, rosulate, simple, pinnatifid, pinnatisect, or runcinate, rarely sinuate or subentire. Cauline leaves petiolate, similar to basal, often less divided upward. Racemes ebracteate, elongated in fruit. Fruiting pedicels slender, rarely thickened, divaricate or erect-ascending. Sepals oblong, ascending, equal, base of lateral pair not saccate. Petals yellow or orange, rarely white or purplish; blade broadly obovate, apex rounded; claw not differentiated from blade. Stamens 6, strongly tetradynamous; filaments of median pairs connate or rarely free, dilated at base; anthers oblong, obtuse at apex. Nectar glands 2, lateral, annular; median glands absent. Ovules 10-40 per ovary. Fruit indehiscent or tardily dehiscent, often lomentaceous siliques, breaking transversely into 1(or 2)-seeded units, linear or oblong, terete, sessile; valves thickened, obscurely veined, glabrous or with glandular and/or dendritic trichomes, torulose or rarely moniliform; replum rounded; septum complete, thickened, opaque, veinless; style prominent or short or obsolete; stigma capitate, strongly 2-lobed, lobes spreading, free, not decurrent. Seeds uniseriate, wingless, oblong, slightly flattened; seed coat not mucilaginous when wetted; cotyledons incumbent.