Shrubs or small trees. Leaves alternate, sessile, small, ericoid, involute, often fasciculate on abbreviated axillary shoots. Capitula small, homogamous, discoid, narrowly cylindric, 1-flowered, in dense clusters or spikes; phyllaries ± 4-seriate, the innermost scarious and longer than the outer; receptacle small, naked. Florets hermaphrodite, narrowly infundibuliform, slightly 5-toothed; anthers sagittate, distal appendage lanceolate; style arms truncate and often penicillate. Achenes glabrous, often 5-ribbed, with a cupuliform apical appendage; pappus of plumose bristles, basally connate or rarely missing.