Biennials or perennials; (short-lived); (glaucous), glabrous throughout. Stems unbranched or branched (several) basally and distally, (0.5-)1.5-12(-15) dm. Basal leaves (soon withered); subrosulate; petiolate; blade broadly ovate, obovate or oblong, 1.5-6 cm, margins entire or repand, or dentate apically. Cauline leaves: blade oblong to obovate or suborbicular, or (distally) orbicular or oblong-ovate, (0.7-)1.5-6(-9) cm × (4-)10-45 mm, base auriculate to amplexicaul (amplexicaul distally), margins entire or repand. Racemes bracteate below or between proximalmost 1 or 2 flowers, (lax to compact, secund or not). Fruiting pedicels divaricate to ascending, (2-)3-12(-17) mm, (expanded at receptacle). Flowers: calyx urceolate; sepals purplish, gray-green, or yellowish, 6-10(-13) mm, keeled or not, (apex recurved); petals purplish or yellowish white (usually with purple veins), (6-)8-14 mm, blade 2-5 × 1-2.5 mm, margins not crisped, claw 4-10 mm, as wide as or wider than blade, (apex reflexed); stamens in 3 unequal pairs; filaments (distinct): abaxial pair (2.5-)4-7 mm, lateral pair (1.5-)3-5 mm, adaxial pair (5-)7-11 mm; anthers (all) fertile or adaxial pair with reduced fertility and shorter, (1.5-)2.5-4.5(-6) mm; gynophore 0.2-1 mm. Fruits arcuate-spreading to pendent, torulose or smooth, straight, flattened, (3-)4-13 (-16) cm × 1.5-2.5(-3) mm; valves each with obscure or somewhat prominent midvein; replum straight; ovules 26-76(-110) per ovary; style 0.4-0.7 mm; stigma entire. Seeds broadly oblong to ovoid or orbicular, 1.5-2.5 × 0.8-1.7 mm; wing 0.1-0.5 mm wide, continuous or on distal 1/2. 2n = 28.