Symbolanthus G.Don

Genus

Angiosperms > Gentianales > Gentianaceae

Characteristics

Herbs, shrubs or rarely small trees; stems usually branched, erect, tetragonal, often slightly winged on the angles. Leaves short petiolate; the petioles amplexi-caul; the lamina generally ovate, elliptic to oblong, subrotund to acuminate at apex, narrow tapering at base, membranous to coriaceous, the costa prominent, the lateral veins becoming obscure. Inflorescences of few-flowered cymes or the flowers solitary, terminal or infrequently axillary. Flowers bracteate, 5-merous, long-pedicellate; calyx campanulate, the lobes free to near the base, carinate; corolla usually pink, sometimes purple, often broadly salverform, the lobes erect to spreading at anthesis, ovate; stamens equalling or exceeding the corolla tube, the filaments inserted on the lower half of the corolla tube, ex-panded basally, the anthers sagittate; ovary ovate, 1-locular, the paired pla-centae parietal along the 2 sutures, the style filiform, the stigma bilamellate, the lobes linear to oblong, flattened. Capsules generally surrounded by the marcescent calyx and corolla, ovate to ellipsoid, septicidally 2-valvate; seeds small, cubical, appearing tuberculate. Pollen grains united in tetrahedral tetrads,
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Herbs, subshrubs, shrubs to small trees. Leaves sessile to petiolate, 1.5-29 cm long; ca. 5-veined. Inflorescences terminal, dichasial with monochasial branches, sometimes flowers solitary; bracts and bracteoles absent/scale-like/leaf-like. Flowers pedicillate, 5-merous, actinomorphic or slightly zygomorphic; calyx fused at base for ca. 1/10 (seldom more) of length, coriaceous, campanulate to tubular; lobes lanceolate, elliptic to ovate, dorsally thickened, often glandular, rarely with keels; apex obtuse to acute; corolla fused at base, salver-shaped to funnel-shaped, 28-110 mm long; corolla bud apex tapering, androecium zygomorphic with stamens and style bent towards bottom of corolla mouth; stamens inserted in corolla tube in lower quarter, of unequal length, base of filaments fused with corolla into a ring-shaped corona, or with flaps of tissue between filament bases, anthers linear, connective apex apiculate; pollen in tetrads; pistil with nectary disk around base, stigma bilamellate. Fruit a capsule, sometimes indehiscent(?), with persistent calyx and deciduous corolla; seeds angular, testa pattern dome-like, sometimes with band-like thickenings.
65-98 microns; single grains 3-colporate (3-colpate, 3-porate), the concurrent colpi of 2 contiguous grains ca. 15 x 6 a, the diameter of apocolpia 25-30 au; exine 4-6 M thick at the distal poles, 7-8 M thick at equator; sexine thicker than nexine, re-ticulate, heterobrochate, differentiated into a coarsely reticulate, ? equatorial zone with lumina 2-20 Mt in diameter, the muri 1-5 M wide, usually supported by one row of bacules, and for the rest a more delicate reticulum with lumina 0.5-5 M in diameter, the muri 1-2 M wide; nexine smooth or granulose (S. pulcher-rimus).
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Foliage retention deciduous
Sexuality hermaphrodite
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Hardiness (USDA) 9-12

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