Syntrichia ruralis (Hedw.) F.Weber & D.Mohr

Great hairy screw-moss (en)

Species

Bryophytes > Pottiales > Pottiaceae > Syntrichia

Characteristics

Plants large, in loose tufts, green to brownish green; terricolous or saxicolous. Stems 5-15 mm tall, branching, tomentose below, tomentum rarely gemmiferous; in section without obvious central strand, inner cortical cells large, thin-walled, outer cortical cells in 1-3 rows, smaller, incrassate, reddish. Leaves appressed, rarely bistratose, mostly contorted dry, recurved to squarrose wet; oblong-spathulate, 2-3 mm long; apex emarginate or truncate, frequently asymmetrical; margins revolute to upper leaf, frequently to apex; base appressed. Costa long-excurrent as hyaline, dentate awn, 2-3 mm long; ventral superficial cells quadrate to short-rectangular, thin-walled, papillose, dorsal superficial cells elongate, incrassate, smooth to rough or spinose; in section guide cells 2, ventral cells in 1-2 layers, large ventral surface cells papillose, dorsal stereid band 4-5 cells thick, reddish, dorsal surface cells generally hyaline, smooth, papillose or spinose. Laminal cells hexagonal, densely papillose, with 4-10 C-shaped papillae over lumen; basal cells large, in well defined group, rectangular, hyaline, smooth; basal marginal cells quadrate, chlorophyllous, papillose. Gemmae round, multicellular, 50-60 µm, green, rarely on tomentum of lower stem or ventral costal surface. Dioicous. Perichaetia terminal, leaves undifferentiated. Seta 10 mm long, reddish brown; capsule cylindrical, 2-3 mm long, brownish; peristome reddish to white, 1.5 mm high, basal membrane long, obliquely tessellated, teeth long-filamentous, papillose; operculum conical, 1 mm long; spores 12-15 µm, finely granular.
More
Stems 5-15 mm. Leaves clasping at base, infolded and twisted around the stem when dry, wide-spreading (in smaller forms) to squarrose-recurved when moist, lingulate-ovate, 1.5-3.5 × 0.75-1.25 mm, canaliculate to keeled; margins tightly revolute in the proximal 7/8 or more, entire; apices emarginate to acute; costa excurrent into a serrate (or occasionally only faintly serrulate), hyaline awn that is often red or sometimes broadly hyaline at base, weakly to strongly papillose on the abaxial surface and often serrate near the apex because of projecting cell ends, red-brown; basal cells abruptly differentiated, narrowly rectangular, 35-70(-90) × 11-18 µm, quadrate to narrowly rectangular at the margins; distal cells quadrate to polygonal, 8-12 µm, with 3-6 papillae per cell, bulging, somewhat obscure. Specialized asexual reproduction absent. Sexual condition dioicous. Seta red, 5-10 mm. Capsule red-brown, 2-3.5 mm, straight, with an abrupt neck; operculum 1.25-1.75 mm, brown; peristome ca. 1.25 mm, the upper divisions twisted ca. 2 turns, red, the basal membrane white, about 1/3 the total length. Spores 11-15 µm, papillose.
Plants large. Stems 5-15 mm long, central strand absent. Leaves wide-spreading to squarrose-recurved when wet, oblong-spathulate, 1.5-3.0 mm long; apex emarginate or truncate; margins revolute to upper leaf; costa long-excurrent as hyaline dentate awn; papillose to spinose on dorsal surface. Upper laminal cells hexagonal, papillose; basal cells abruptly differentiated, narrowly rectangular, hyaline. Capsule straight, peristome teeth twisted.
Life form -
Growth form -
Growth support -
Foliage retention deciduous
Sexuality -
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.01 - 0.02
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity 4-6
Soil texture 7-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) -

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Syntrichia ruralis unspecified picture

Distribution

Syntrichia ruralis world distribution map, present in United States of America and South Africa

Identifiers

LSID 35177789
WFO ID wfo-0001191362
COL ID 53ZRQ
BDTFX ID -
INPN ID 434242
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Tortula ruraliformis Tortula ruralis Tortula longimucronata Syntrichia ruraliformis Syntrichia ruralis Barbula ruraliformis Barbula ruralis Tortula ruralis var. ruraliformis Barbula ruralis var. ruraliformis Syntrichia ruralis var. glacialis

Lower taxons

Syntrichia ruralis var. densiretis Syntrichia ruralis var. epilosa Syntrichia ruralis var. gigantea Syntrichia ruralis var. gracilis Syntrichia ruralis var. subpapillosissima