Deciduous, buttressed tree. Leaf scars prominent, conspicuous. Leaves roundish, acuminate, entire or dentate, undersurface densely hairy by bulbous-based simple hairs, upper surface subglabrous or sparsely set with simple hairs. Flowers of both sexes subsessile or shortly pedicelled, in fasciculate apical, peduncled, pendent, simple or little branched spikes (♀) or panicles (♂) with tomentose rhachis on the apex of thick defoliate twigs, solitary or in twos or threes, 4-(6) or 4-5-merous (♀) ♂ Flowers: calyx tube short, lobes oblong to ovate, equal, or inequal, sometimes alternating with a few occasional smaller, narrower appendages in structure similar to the calyx lobes. Petals 0. Stamens 4, opposite the sepals, inserted on the margin of the cup-shaped receptacle. Filaments short or rather long, anthers incurved in bud, 2-celled, extrors ± basifix, at last seemingly peltate. Rudimentary ovary disk-shaped, cross-like, or 0. ♀ Flowers: lower part of the calyx connate with the ovary, slightly 4-angular, glandular, and mostly hairy, upper part cupular, with 4 triangular lobes. Petals and androecium 0. Styles 4-5, erect, opposite the calyx lobes, inserted on the margin of the calyx throat, thickish, persistent, with an obliquely inserted, unilaterally stigmatic, thickened apex. Ovary with 4-5 parietal placentas alternating with the styles. Ovules in several rows. Flat top of the capsule splitting before the styles, the thus formed 4 triangular valves marcescent and curving inwards, leaving a roundish apical pore. Seeds narrow-oblong, slightly flattened.
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Trees deciduous, buttressed. Male flowers 4-or 5-conglomerate on spikes; calyx deeply 4(or 5)-lobed, tube very short, cupular; petals absent; stamens 4(or 5); filaments longer than calyx lobes, incurved with extrorse anthers in bud, erect with introrse anthers at anthesis; sterile ovary discoid-subcruciate, rarely absent. Female flowers solitary or 2-4-conglomerate on spikes; calyx tube long, slightly quadrangular, cupular in distal part, lobes 4(or 5), triangular; ovary with 4(or 5) parietal placentas; styles 4(or 5). Capsule dehiscing apically between persistent styles. Seeds ovoid.
Dioecious, deciduous, buttressed trees. Leaves simple, long-petiolate, dentate to almost entire, hairy or tomentose below. Flowers in elongate slender spikes, racemes or panicles, produced before leaves. Male flowers: calyx tube short, 4-lobed; petals absent; stamens 4. Female flowers: calyx tube 4 dentate; petals absent; stamens absent; styles 4, rarely 5; stigmas slightly dilated. Capsule dehiscing at apex between styles. Seeds narrowly oblong, slightly flattened.
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