Toddalia asiatica (L.) Lam.

Species

Angiosperms > Sapindales > Rutaceae > Toddalia

Characteristics

Sarmentose or climbing shrubs, retrorsely aculeolate; young branches and leaves ferruginous-pubescent, glabrescent. Leaves with petiole 1-4 cm long, grooved above and persistently pubescent in the groove, sometimes aculeolate; leaflets sessile; lamina 3-7(8) x 1.2-3 cm, elliptic or obovate or oblanceolate, that of the lateral leaflets asymmetric, chartaceous, conspicuously glandular on both surfaces, apex shortly acuminate or obtuse, margin entire or slightly crenulate; midrib prominent on the lower surface and occasionally aculeolate; lateral nerves and veins numerous, not raised. Inflorescence shorter or as long as the subtending leaf, ferruginous-pubescent; bracts 0.8-1.3 mm long, linear, caducous. Flowers 5-merous, unisexual by abortion, usually in cymose clusters on the panicle-branches; pedicels 1-3 mm long, acute, caducous. Petals 2.5 mm long, valvate. Male flowers: stamens as long as the petals; anthers 1 mm long, 2-lobed, ovate. Female flowers not seen. Fruit 7-8 mm in diam., drupaceous, 3-5(7)-locular. Seeds 4 mm long, reniform; testa dark brown, hard, smooth; endosperm fleshy.
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A scrambling or climbing, retrorsely aculeolate shrub; young branches minutely rusty pubescent or glabrescent.. Leaflets sessile, elliptic or obovate or oblanceolate, 3–8 cm. long, 1.3–3 cm. wide, acuminate or obtuse at the apex, entire or slightly crenulate; midrib prominent and occasionally aculeolate on the lower surface; lateral nerves numerous but not raised; conspicuously glandular on both surfaces; petiole 1–4 cm. long, grooved above and persistently pubescent in the groove, sometimes aculeolate.. Inflorescence ferruginous-pubescent; bracts linear, 0.8–1.3 mm. long, caducous.. Flowers usually in cymose clusters on the panicle branches, unisexual by abortion; pedicels 1–3 mm. long.. Calyx small, with 5 acute lobes.. Petals triangular-lanceolate, ± 2.5 mm. long, yellow.. Male flowers with ovate anthers 1 mm. long; rudimentary ovary glabrous, with a long 5-ribbed style.. Female flowers with a very short style and 3–5-lobed stigma; 5 staminodes present.. Fruit 7–10 mm. across, orange when ripe.. Seeds 3–4 mm. long, dark brown, smooth.. Fig 1.
Shrubs (usually sprawling) or woody climbers, usually armed. Petiole 1-4 cm; leaflet blades usually sessile or subsessile, elliptic or narrowly elliptic to obovate to oblanceolate, 3-10 × 1-4 cm, base narrowly cuneate to attenuate, apex acuminate or rarely acute to obtuse or rounded. Inflorescences to 17 cm. Sepals 0.3-0.5 mm. Petals cream-white, ovate to elliptic, 1-3.5 mm. Stamens in male flowers 3-4 mm, in female flowers ligulate and 0.2-0.8 mm. Disk 0.2-0.5 mm. Gynoecium in female flowers ovoid to ellipsoid and 1.5-2.5 mm, in male flowers subcylindric and 1-2 mm. Fruit 5-10 mm in diam. Seeds 5-6.5 mm. Fl. year-round but mostly in spring and summer, fr. autumn and winter.
A spiny and woody vine. It is a shrub that grows 2-20 m long. The prickles are curved. The leaves are alternate and have 3 leaflets. The leaflets are pointed at both ends. The leaf stalk is 2 cm long. The leaflets are 3-10 cm long by 1-2.5 cm wide. The flowers are small, greenish and borne in large compound flower arrangements. This is 7.5 cm long. Male and female flowers are separate. The fruit are small and borne in fairly large clusters. They can be 3 to 5 angled. The fruit are 5-7 mm across. There are up to 7 seeds.
Leaves with petiole 1–4 cm. long, grooved above and persistently pubescent in the groove, sometimes aculeolate; leaflets sessile; lamina 3–7 (8) × 1·2–3 cm., elliptic or obovate or oblanceolate, that of the lateral leaflets asymmetric, chartaceous, conspicuously glandular on both surfaces, apex shortly acuminate or obtuse, margin entire or slightly crenulate; midrib prominent on the lower surface and occasionally aculeolate; lateral nerves and veins numerous, not raised.
Climber or scrambler. Branches with small recurved prickles. Leaves 3-foliolate, petiole grooved above, leaflets with midribs prominent on lower surface and occasionally aculeolate. Flowers yellow.
Flowers 5-merous, unisexual by abortion, usually in cymose clusters on the panicle-branches; pedicels 1–3 mm. long, acute, caducous.
Inflorescence shorter or as long as the subtending leaf, ferruginous-pubescent; bracts 0·8–1·3 mm. long, linear, caducous.
Sarmentose or climbing shrubs, retrorsely aculeolate; young branches and leaves ferruginous-pubescent, glabrescent.
Male flowers: stamens as long as the petals; anthers 1 mm. long, 2-lobed, ovate.
Seeds 4 mm. long, reniform; testa dark brown, hard, smooth; endosperm fleshy.
Fruit 7–8 mm. in diam., drupaceous, 3–5 (7)-locular.
Petals 2·5 mm. long, valvate.
Female flowers not seen.
Life form perennial
Growth form shrub
Growth support climber
Foliage retention evergreen
Sexuality -
Pollination entomogamy
Spread -
Mature width (meter) -
Mature height (meter) 12.5
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a tropical plant. Common throughout the islands of the Philippines in secondary forest and virgin forest at low altitudes but up to 1700 m. In Yunnan in China it grows up to 1500 m above sea level. In Swaziland it grows in the low veld only. It grows in the lowlands. In Sichuan. It occurs in the Western Ghats in India.
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Forests near rivers or streams. Thickets and forests near coasts, especially in calcareous regions, in southern Japan. Humid forests, from sea-level up to 2300 metres.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

The fruit are used for food flavouring. They have a pungency like pepper. They are crushed. All parts of the plant including the leaves are cooked and can be used for flavouring. The fruit are boiled to make soup. The ripe fruit are pickled.
Uses dye essential oil fodder food gene source material medicinal
Edible fruits leaves
Therapeutic use Insect bites and stings (flower), Antifungal agents (leaf), Colic (leaf), Parasympatholytics (leaf), Antifungal agents (root), Antipyretics (root), Appetite stimulants (root), Asthenia (root), Bronchitis (root), Colic (root), Convalescence (root), Cough (root), Diarrhea (root), Dyspepsia (root), Epilepsy (root), Expectorants (root), Fever (root), Flatulence (root), Gonorrhea (root), Malaria (root), Nausea (root), Paralysis (root), General tonic for rejuvenation (root), Stomatitis (root), Toothache (root), Ulcer (root), Wound healing (root), Wounds and injuries (root), Antiperiodic (root), Diaphoretic (root), Bactericide (unspecified), Carminative (unspecified), Cough (unspecified), Debility (unspecified), Diaphoretic (unspecified), Diarrhea (unspecified), Flu (unspecified), Gonorrhea (unspecified), Malaria (unspecified), Poison (unspecified), Rheumatism (unspecified), Stimulant (unspecified), Sting(Wasp) (unspecified), Stomachic (unspecified), Amenorrhea (unspecified), Fever (unspecified), Stomach (unspecified), Tonic (unspecified), Antipyretics (unspecified), Asthenia (unspecified), Parasympatholytics (unspecified), General tonic for rejuvenation (unspecified), Stomatitis (unspecified), Anthelmintics (whole plant), Antipyretics (whole plant), Antirheumatic agents (whole plant), Diarrhea (whole plant), Digestive system diseases (whole plant), Diuretics (whole plant), Fever (whole plant), Leprosy, lepromatous (whole plant), Pain (whole plant), Parasympatholytics (whole plant), Antiperiodic (whole plant), Diaphoretic (whole plant)
Human toxicity -
Animal toxicity -

Cultivation

Plants can be grown from seeds.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Toddalia asiatica habit picture by E. Mouysset (cc-by-sa)

Leaf

Toddalia asiatica leaf picture by E. Mouysset (cc-by-sa)
Toddalia asiatica leaf picture by E. Mouysset (cc-by-sa)
Toddalia asiatica leaf picture by E. Mouysset (cc-by-sa)

Flower

Toddalia asiatica flower picture by Bernard REYNAUD (cc-by-sa)

Distribution

Toddalia asiatica world distribution map, present in Bangladesh, Bhutan, China, Algeria, Indonesia, Sri Lanka, Myanmar, Nepal, Philippines, Thailand, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:775412-1
WFO ID wfo-0000455337
COL ID 7CG53
BDTFX ID -
INPN ID 706870
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Aralia labordei Toddalia willdenowii Toddalia effusa Toddalia ambigua Toddalia aculeata Toddalia rubicaulis Toddalia micrantha Paullinia asiatica Limonia oligandra Cranzia nitida Cranzia schmidelioides Cranzia willdenowii Toddalia schmidelioides Toddalia floribunda Toddalia rubricaulis Toddalia tonkinensis Toddalia angustifolia Toddalia nitida Cranzia aculeata Cranzia asiatica Toddalia asiatica var. floribunda Toddalia asiatica var. gracilis Toddalia asiatica var. obtusifolia Toddalia asiatica