Toddalia Juss.

Genus

Angiosperms > Sapindales > Rutaceae

Characteristics

Shrubs (usually sprawling) or woody climbers, dioecious, usually armed. Leaves alternate, digitately 3-foliolate (occasional leaves 1-or 2-foliolate). Inflorescences terminal and/or axillary, paniculate, racemose, or umbelliform. Sepals 4 or 5(or 6), connate at base or to ± half their length. Petals 4 or 5(or 6), valvate or narrowly imbricate in bud. Stamens 4 or 5(or 6), distinct, rudimentary in female flowers. Disk pulvinate. Gynoecium 4-7-loculed, syncarpous, rudimentary in male flowers; ovules 2 per locule; stigma sessile or nearly so. Fruit a 4-7-loculed drupaceous berry; exocarp fleshy; mesocarp undifferentiated; endocarp cartilaginous. Seeds brown to black, reniform, dull to lustrous; seed coat with thick inner layer of dense black sclerenchyma surrounded by outer layer of compact parenchymatous tissue; endosperm copious; embryo curved; cotyledons elliptic, flattened; hypocotyl superior.
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Woody scramblers; branches usually armed with prickles. Leaves trifoliolate, alternate, sometimes aculeate. Inflorescence of axillary and terminal panicles. Flowers unisexual by abortion, 5-merous, small. Sepals united to form a cupuliform 5-dentate calyx. Petals 5, free, imbricate. Male flowers: stamens 5, inserted at the base of a short gynophore, opposite the sepals; filaments filiform; ovary vestigial. Female flowers: staminodes filiform; ovary on a short gynophore, 5–7-locular, with 2 ovules in each locule; stigma 3–5-lobed, disk-shaped. Fruit globose, drupaceous, with woody endocarp, 3–7-locular, locule 1-seeded. Seed oblong-reniform; testa thick; cotyledons linear.
Male flowers: stamens 5, inserted at the base of a short gynophore, opposite the sepals; filaments filiform; anthers introrse; ovary vestigial, conical.
Female flowers: staminodes filiform; ovary on a short gynophore, 5–7-locular, with 2 ovules in each loculus; stigma disk-shaped, 5-lobed.
Sarmentose or climbing shrubs; branches, leaves and inflorescences aculeate.
Fruit globose, drupaceous with woody endocarp, 3–7-locular; loculi 1-seeded.
Seeds reniform, oblong; testa thick; cotyledons linear.
Sepals united to form a cupuliforrn 5-dentate calyx.
Inflorescence of axillary and terminal panicles.
Flowers small, 5-merous, unisexual by abortion.
Leaves alternate, 3-fbliolate.
Petals 5, free, imbricate.
Life form perennial
Growth form shrub
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Cultivation

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Images

Toddalia unspecified picture

Distribution

Toddalia world distribution map, present in China

Identifiers

LSID urn:lsid:ipni.org:names:329424-2
WFO ID wfo-4000038512
COL ID 7NDP3
BDTFX ID -
INPN ID 705342
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Duncania Toddalia

Lower taxons

Toddalia asiatica