Stem rounded-pentagonal in section. Leaves flattened, ovate to elliptical, distal margins plane or weakly erect or rarely broadly involute, entire, not bordered; apex rounded or rounded-acute, plane to weakly concave, keeled or cucullate; basal cells evenly differentiated across leaf base or as a U, only weakly running up margins if at all, not distinctly enlarged submarginally; distal laminal cells pluripapillose with low papillae; mucro short-conic, of 3-4 cells. Sexual condition autoicous. Peristome teeth absent to short-lanceolate.