Plants turf-forming, yellowish green distally, medium brown to light brown basally. Stems to ca. 3 cm, hyalodermis present or occasionally absent, sclerodermis usually weak, often of substereid cells, central strand present or absent; axillary hairs with proximal 1-3 cells thicker walled or all hyaline. Leaves incurved, often catenulate, often reflexed at top of appressed base when dry, weakly to widely spreading when moist; oblong, elliptical or ligulate to long-lanceolate, adaxial surface flat or shallowly channeled, 1.5-2.5(-5) mm; base elliptic to rectangular, occasionally not differentiated in shape; distal margins plane or somewhat erect but never sharply incurved, entire to crenulate-notched or occasionally dentate in the distal 1/2-3/4, occasionally with a narrow, less papillose border; apex narrowly to broadly acute or rounded, occasionally broken or reflexed; costa usually excurrent as a smooth, sharp mucro, adaxial outgrowths absent, adaxial cells quadrate but occasionally elongate near apex, in 2-6(-8) rows; cross section semicircular, reniform or ovate, adaxial epidermis present, adaxial stereid band present, occasionally weak, guide cells 2-4(-6) in 1 layer, hydroid strand usually absent, abaxial stereid band present, crescentlike in section, abaxial epidermis weak or absent; basal cells differentiated across leaf or medially, occasionally rising weakly along margin, rectangular, seldom bulging, 3-5:1; distal cells rounded-quadrate, occasionally transversely elliptical along margins, 6-12(-18) µm wide, papillae usually 2-fid, crowded, 2-6 per lumen, occasionally single and multiplex and covering the lumens, superficial cell walls equally thickened and convex on both sides. Specialized asexual reproduction absent [rare, of gemmae on rhizoids or adaxial surface of costa, of several cells, vermiform to irregular in shape, occasionally branching]. Sexual condition dioicous or occasionally autoicous. Perichaetia terminal, interior leaves weakly sheathing at base or not sheathing, little different from cauline leaves. Capsule stegocarpic [cleistocarpic]. Seta 0.4-1.5 cm. Theca cylindric, ovate or elliptic ca. 1-3 mm, annulus of 1-4 rows of vesiculose cells, persistent or seldom revoluble; operculum [when differentiated] long-conic to rostrate; peristome teeth 16, usually rather short, occasionally rudimentary or absent, ligulate to filamentous, entire or occasionally irregularly cleft 2-3-fid or perforate, straight or seldom weakly twisted counterclockwise. Calyptra cucullate. Spores 8-25 µm. KOH laminal color reaction medium orange to yellowish orange, occasionally yellow.
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