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Trees or shrubs, monoecious. Leaves simple, alternate, rarely opposite to nearly whorled; stipules small, persistent or deciduous; leaf blade unlobed, pinnately veined, sometimes stipulate at base, paired glands at apex of petiole. Flowers androgynous or anisogynous; inflorescences terminal or axillary, racemose or thyrsoid. Male flowers: sepals 5, free, imbricate; petals 5, exceeding calyx; disk cupular or annular, or shortly divided, or dissected, sometimes divided into 5 glands; stamens 3-5; filaments connate or only free at upper part; anther connective enlarged, often elongated, thecae adnate to thickened connectives; pistillode absent. Female flowers: pedicels usually prominently stronger than axil of inflorescence; sepals and petals as in male; disk annular or cupular, usually unlobed; ovary 3-locular; ovules 1 per locule; styles 3, free, connate at near base, bifid or unlobed at apex. Capsules 3-valved. Seeds not carunculate; testa dry; endosperm present; endosperm fleshy; cotyledon broad and flat.
