Trees , to 30 m; crowns oblong. Bark gray, deeply fissured with broad, flattened ridges. Wood hard. Branches short-spreading, young branches pubescent, old-growth with 3-5 prominent, irregular, corky wings; twigs reddish, pubescent. Buds brown, ovoid, acute, pubescent; scales brown, pilose on outer surface, ciliate on margins. Leaves: petiole ca. 5 mm, pubescent. Leaf blade obovate to oblong-oval, (2.5-)9-11(-16) × 2.5-5 cm, base oblique, margins doubly serrate, apex short-acuminate; surfaces abaxially white-pubescent, pubescence not tufted in axils of veins, adaxially dark green, usually glabrous, sometimes scabrous. Inflorescences racemose cymes, long-pendulous, (7-)10(-13)-flowered, to 5 cm; pedicel 0.5-1 cm. Flowers: calyx deeply lobed, divided nearly to middle, lobes 7-8; stamens 5-8; anthers dark purple; stigmas greenish, pubescent. Samaras elliptic to oval, 1.5-2.2 cm, narrowly winged, pubescent, margins short-ciliate, apex shallowly notched. Seeds inflated, not thickened. 2 n = 28.
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A medium sized tree. It grows to 25-30 m high. It spreads 12 m wide. The trunk is 75 cm across. It is an upright tree with a narrow rounded crown. The young branches have corky bark. The leaves are 5-10 cm long and 2-5 cm wide. They have teeth along the edge. The leaves are shiny dark green above and pale green with soft hairs underneath. The flowers are 3 mm wide and green. They occur in long drooping clusters. The fruit are 10-19 mm long and flat keys. They have one seed and wide wings. These are notched at the tip.
Tree to 30 m, the thinly hairy twigs often becoming irregularly winged with 2 or more plates of cork after their second year; buds thinly hairy; lvs oblong to obovate, 8–14 cm, distinctly cordate on one side at base, glabrous above; fls in slender racemes to 4 cm; cal merely lobed; fr elliptic, 1.5–2.3 cm, pubescent on the sides, the stipe 1 mm; 2n=28. Rich upland woods; sw. Que. to Minn., s. to N.J., W.Va., w. Va., Tenn., and n. Ark. (U. racemosa Thomas)
Rich woods and calcareous uplands. Dry gravelly uplands, low heavy clay soils, rocky slopes and river cliffs. Rocky slopes, limestone outcrops, rich woods, flood plains, stream banks; at elevations from 30-900 metres.
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It often grows on heavy clay soils near limestone ridges. It grows between 60-760 m altitude. It suits hardiness zones 2-9.
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