Verbena litoralis Kunth

Seashore vervain (en), Verveine (fr)

Species

Angiosperms > Lamiales > Verbenaceae > Verbena

Characteristics

Quick-growing erect or suberect perennial herbs, 0.2-2 m high, sometimes large and robust, woody-based, fastigiately few-to many-branched above, glabrous and shiny or sparingly strigillose; stems to 12 mm thick, tetragonal or subtetragonal in cross-section, the internodes deeply sulcate above the insertion of the leaves and faintly many-sulcate between, scabridulous on the angles when young, green, slightly contracted at the nodes. Leaves with the blades lanceolate to oblong or oblanceolate, 3-10 cm long and 1-1.5 cm wide, acute apically, tapering into a short petiole basally or sessile to subsessile, acuminate and somewhat clasping basally, scabridulous throughout or scabrous and somewhat rugose only above, the upper 2/3 more or less sharply and coarsely serrate with commonly 3-5 some-what mucronate teeth per side, sparsely strigillose ,on both surfaces, the venation impressed above and prominent beneath. Inflorescences terminal spikes, several to many, pedunculate, cymose or corymbose-paniculate, slender or filiform, pilo-sulous, at first congested and about 4 mm in diameter, dense or interrupted, later loosely flowered and elongate, in fruit becoming 2-8 cm long, the fruits usually not contiguous; bractlets ovate-lanceolate, marcescent, rigid, acuminate or sub-ulate, 1-3 mm long, usually subequaling or somewhat shorter than the calyx, abruptly upcurved, glabrate. Flowers with the calyx often purple, 2-2.5 mm long, pubescent or finely strigillose, the rim subtruncate, the 5 teeth equal, minute, subulate; corolla ca. 3 mm long, various shades of blue, violet, or purple, puberu-lent, the tube greenish toward the base, variable in length, always somewhat longer than the calyx, the limb inconspicuous, 2.5-3 mm wide, puberulent on the inner surface, the lobes subequal, obtuse, glabrous; anthers greenish-yellow; ovary almost 1 mm long, green, glabrous, the style ca. 1 mm long. Fruits with the nutlets trigonous, dark-stramineous, linear-oblong, 1-1.9 mm long, glabrous, striate, some-what reticulate, the commissural face about as long as the nutlet, muricate-scabrous.
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Short-lived perennial; stems square, somewhat scabrid, to c. 1 m tall. Lvs mostly petiolate, the uppermost sometimes subsessile. Lamina of lower lvs to c. 10 × 2.5 cm, lanceolate to oblong or rhomboid, with ± strigulose hairs scarcely swollen at base above and below, usually coarsely or deeply serrate; veins not impressed above; base attenuate; apex acute. Infl. loosely paniculate; spikes to c. 5 cm long at maximum flowering, hairy, elongating to c. 15 cm long at fruiting, slender; fls rather dense but soon becoming distant. Bracts ± = calyx at flowering, lanceolate, keeled, acuminate, ciliate. Calyx 2-3 mm long, glandular-hairy; teeth green, acute. Corolla tube > calyx, rather sparingly hairy outside; limb 2-3 mm diam., bluish or mauve, drying a similar colour. Nutlets c. 1.5 mm long, oblong, faintly ribbed dorsally, brown, finely white-papillate on flattened ventral surface.
Perennial herb or shrub, 1-2 m high; stem erect or suberect, branched above, 4-angled, with whitish angles and flat faces. Leaves ovate, rhombic-ovate, spathulate or oblanceolate, 20-50 x 10-20 mm, apex acute, base tapering into narrow petiole, margins serrate to double serrate, teeth sometimes obtuse; both surfaces scabrid with sparse, short-appressed, antrorse, tubercle-based hairs, bristles more numerous on lower surface. Inflorescences much branched, with several to many corymbose-paniculately arranged spikes; spikes slender or filiform; bracts lanceolate. Flowers lavender or lilac to violet or violet-blue. Calyx hirsute, hispidulous or tomentose. Corolla: tube exserted from calyx; limb inconspicuous, with subequal, obtuse lobes. Fruit dry, enclosed in calyx, separating at maturity into four 1-seeded, crustaceous nutlets.
Herb 0.3–1 m high; branches erect, quadrangular, smooth, sparsely hairy (eglandular). Leaves shortly petiolate or sessile; lower leaves variously acutely serrate, 1-or 2-pinnatifid, 2–7 cm long, 0.5–3 cm wide; upper leaves incised to subentire, smaller than lower leaves, usually ±narrowly ovate; base ±acute; apex acute (for both upper and lower leaves). Inflorescence a lax terminal panicle with long branches ending in a spike; bracts at least reaching top of calyx, with peduncle, bracts and calyx strongly glandular. Calyx 5-toothed, 2.5–3 mm long. Corolla pale pink or lilac, slightly longer than calyx; tube c. 2.5 mm long. Mericarps 1.5–2 mm long.
Perennial herb or shrub, up to 2 m high. Stems erect or suberect; 4-angled. Leaves simple, petiolate or sessile, not amplexicaule; blade ovate, rhombic ovate or narrowly obovate, 20-50 x 10-20 mm, margins serrate to double serrate, base attenuate. Flowers: in much-branched inflorescences with lax groups of triads; triads with median spike usually longer and subsessile, lateral ones on peduncles, 5-8 mm long; calyx 2-3 mm long; corolla 3.0-3.5 mm long; corolla violet or purple; Nov.-May.
Herb, up to 0.8 m high. Leaves petiolate or tapering to cuneate-attenuate base, 30-100 mm long, lanceolate or oblong-lanceolate, scabrid. Inflorescence 750 mm long, dense at apex, more open below and finely strigillose. Flowers purple.
A herb. It keeps growing from year to year. The leaves are 5-12 cm long by 1-3 cm wide.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination entomogamy
Spread -
Mature width (meter) -
Mature height (meter) 1.0
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Wet or dry thickets, meadows, rocky slopes, often a weed in cultivated ground, sometimes in pine-oak forest; at elevations from 85-3,000 metres.
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Wet or dry thickets, meadows, rocky slopes, often a weed in cultivated ground, sometimes in pine-oak forest; at elevations from 85-3,000 metres.
It is a subtropical plant. In Argentina it grows from sea level to 2,000 m above sea level.
Grows in wastelands and open areas along watercourses, roads and tracks.
Light -
Soil humidity 6-8
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

The leaves are used for tea.
Uses medicinal tea
Edible leaves
Therapeutic use Fertility agents (leaf), Fertility agents (stem), Antifertility (unspecified), Cough (unspecified), Panacea (unspecified), Purgative (unspecified), Tumor (unspecified), Malaria (unspecified), Bruise (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings. Seeds needs stratification.
Mode seedlings
Germination duration (days) 14 - 90
Germination temperacture (C°) 18
Germination luminosity dark
Germination treatment stratification
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Verbena litoralis leaf picture by Siesquen Juan Carlos (cc-by-sa)
Verbena litoralis leaf picture by Kiera O’Halloran (cc-by-sa)

Flower

Verbena litoralis flower picture by Jimi Dias Maggi (cc-by-sa)
Verbena litoralis flower picture by Génesis Monroy (cc-by-sa)
Verbena litoralis flower picture by A M (cc-by-sa)

Distribution

Verbena litoralis world distribution map, present in Australia, Belize, Bolivia (Plurinational State of), Brazil, Costa Rica, Ecuador, Guatemala, Honduras, Mauritius, Norfolk Island, Nicaragua, New Zealand, Panama, Peru, Puerto Rico, Paraguay, Réunion, Uruguay, United States of America, Venezuela (Bolivarian Republic of), South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:30007120-2
WFO ID wfo-0000332038
COL ID 5B2RT
BDTFX ID 82380
INPN ID 446084
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Verbena longifolia f. albiflora Verbena affinis Verbena caracasana Verbena lanceolata Verbena sedula Verbena gentryi Verbena longifolia Verbena nudiflora Verbena paucifolia Verbena integrifolia f. albiflora Verbena litoralis f. litoralis Verbena integrifolia Verbena brasiliensis var. subglabrata Verbena carolina var. glabra Verbena glabrata var. tenuispicata Verbena litoralis var. albiflora Verbena litoralis var. caracasana Verbena litoralis var. leptostachya Verbena litoralis var. portoricensis Verbena litoralis var. pycnostachya Verbena longifolia var. pubescens Verbena minutiflora var. peruviana Verbena sedula var. fournieri Verbena litoralis var. melanopotamica Verbena bonariensis var. litoralis Verbena litoralis var. glabrior Verbena sedula var. darwinii Verbena litoralis f. albiflora Verbena litoralis f. magnifolia Verbena litoralis var. litoralis Verbena litoralis var. subglabrata (moldenke) n.o'leary Verbena litoralis