Verbena L.

Vervain (en), Verveine (fr)

Genus

Angiosperms > Lamiales > Verbenaceae

Characteristics

Herbs, sometimes slightly woody basally; stems and branches procumbent, 3-scending, or erect, glabrous or variously pubescent. Leaves mostly decussate-Dpposite, rarely whorled, dentate (rarely entire) or variously lobed, incised, or pinnatifid. Inflorescences spicate; spikes terminal, usually densely many-flowered, Dften flat-topped and pseudo-umbellate or fasciculate-capitate, sometimes greatly elongate with densely crowded or scattered flowers, rarely also axillary, often greatly elongating only after anthesis. Flowers small or medium-sized, each solitary and axillary in a usually narrow bractlet; calyx usually tubular, 5-angled, 5-ribbed, unequally 5-toothed, not at all or but slightly changed in fruit, then herbaceous, mostly ventricose, eventually split, often connivent or contorted apically, thin-textured; corolla salverform or-funnelform, the tube cylindric, straight or curved, often slightly ampliate or subinflated apically, usually villous at the stamen-insertion within and barbate at the mouth, the limb flat, spreading or oblique, weakly 2-lipped, the lobes 5, usually elongate, more or less unequal, varying from obtuse or rounded to emarginate apically, the 2 posterior outermost and the anterior innermost in prefloration; stamens 4, didynamous, inserted in the upper half of the corolla-tube, included, the anthers ovate, with 2 parallel or slightly divergent thecae, dehiscing longitudinally, the dorsal connective un-appendaged and narrowly muticous or provided with a more or less conspicuous glandular appendage; gynoecium 2-carpelled, the ovary superior, entire or 4-lobed, apically situated on a somewhat lobed annular disc, completely 4-loculed even at anthesis, the ovules 1 per locule, attached laterally at or near the base, erect, anatropous; the style usually short and equaling the stamens, somewhat dilated, shortly 2-lobed or bifid apically, the anterior lobe or branch stigmatiferous and subpulvinate-papillose, the posterior lobe or branch smooth and horny and non-stigmatiferous, not as broad as the anterior. Fruit mostly enclosed by the mature calyx, schizocarpous, the pericarp hard and dry, readily separating at maturity into 4, 1-seeded linear or linear-oblong (rarely elliptic-winged) crustaceous nut-lets; seeds 4, erect, oblong, triangular in cross-section, without or with negligible endosperm, the outer face convex, the cotyledons 2, thick, applicate.
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Herbs, sometimes slightly woody at base, or subshrubs, open or decumbent, matted or tufted, rarely glabrous; branches quadrangular or almost so. Leaves mostly decussate, rarely whorled, small, entire or variously incised or pinnatifid. Inflorescence usually much branched, sometimes a capitate spike (when young), often elongating after flowering, terminal, rarely axillary, usually densely many-flowered. Flowers solitary in axil of a minute, inconspicuous bract. Calyx usually tubular, unequally and minutely 5-toothed, or only 4-toothed (V. supina); teeth not contorted at maturity. Corolla funneliform; tube straight or slightly curved; lobes 5, slightly unequal and spreading. Stamens 4, didynamous, included in corolla tube. Ovary with 2 carpels, each carpel 2-locular, hence ovary appearing 4-locular with 1 ovule per loculus; style less than 3 times length of ovary. Fruit dry, readily separating at maturity into 4, 1-seeded mericarps; pericarp not extended basally.
Annual or perennial herbs or subshrubs. Leaves opposite or sometimes in whorls of 3, simple and dentate or deeply divided, rarely entire. Flowers in simple spikes or panicles of spikes, often dense and compact or more elongate and lax, bracteate, terminal. Calyx tubular, distinctly and ± unequally (4–)5-toothed, 5-ribbed. Corolla funnel-shaped or hypocrateriform; tube straight or ± curved, ± pubescent within; limb 5-lobed, ± spreading, feebly to distincdy 2-lipped. Stamens (2–)4(–5), included, usually inserted near the middle of the tube but sometimes higher, didynamous. Ovary entire or 4-lobed, 4-locular with a single ovule in each locule; style slender but ± short; stigma unequally 2-lobed. Fruit splitting into 4 nutlets at maturity, enveloped in the persistent calyx. Nutlets with inner face often scabridulous or granulate.
Annual or perennial, taprooted or rhizomatous herbs or subshrubs, unarmed, usually hairy. Lvs usually opposite, rarely in whorls of 3 or alternate, usually toothed or lobed. Infl. usually of 1 or more terminal spikes, these often corymbosely or paniculately arranged; fls sessile or subsessile, usually rather small, bracteate. Calyx tubular, 5-ribbed and unequally 5-toothed, ± accrescent. Corolla salverform; tube straight or curved, usually > calyx; limb 5-lobed and ± 2-lipped, the lobes obtuse or emarginate. Stamens 4, didynamous, included. Style included. Ovary 4-celled; ovules 1 to each cell. Fr. dry, separating into 4 nutlets.
Fls in spikes (or heads) ending the stem and branches; cal tubular, 5-angled and unequally 5-toothed, persistent, scarcely changing in fr; cor salverform or funnelform, the tube straight or curved, the limb flat, weakly 2-lipped, with 5 obtuse or emarginate lobes; stamens 4, paired, included; anther-connective often with a glandular appendage; style terminal, shortly 2-lobed, only one lobe stigmatic; ovary with 4 uniovulate chambers, often rather shallowly 4-lobed, separating at maturity into 4 nutlets, these often ridged or reticulate; ours herbs. (Glandularia) 250, mostly New World.
Herbs or subshrubs, annual or perennial. Branches 4-veined. Leaves opposite, sessile, dentate, lobed, or pinnatifid. Spikes usually terminal, simple or branched; bracts small. Flowers alternate, nearly zygomorphic. Calyx membranous, 5-dentate. Corolla with a distinct tube; lobes 5, unequal, spreading. Stamens 4, fertile, didynamous, inserted at middle of corolla tube, included; anthers ovate. Ovary 4-locular; ovules erect, 1 per locule. Capsules dry, included in calyx, dehiscing into 4 1-seeded pyrenes.
Corolla funnel-shaped or hypocrateriform; tube straight or curved, cylindric or widening slightly towards the apex, usually villous within particularly at the mouth and about the insertion of the filaments; limb spreading, oblique, weakly 2-lipped, with unequal obtuse lobes, rounded or emarginate at the apex.
Ovary 2-carpellate, entire or slightly lobed at the apex, 4-locular at anthesis, each locule l-ovulate; ovules anatropous, erect, attached laterally at or near the base; style usually equalling the stamens, divided at the apex into a short, stigmatic anterior lobe and an acute horny sterile posterior tooth.
Stamens 4, included, didynamous; filaments inserted about or above the middle of the corolla tube; anthers ovate, with parallel or diverging thecae without appendages, or the connectives of upper 2 stamens each provided with a clavate or glanduliform appendage.
Fruit a schizocarp, enclosed within the persistent calyx, separating when ripe into four 1-seeded mericarps; mericarps linear or oblong, with dry pericarp, longitudinally ribbed with a raised reticulation toward the apex.
Inflorescences spicate with the spikes terminal, sometimes axillary, simple or corymbosely or paniculately arranged; spikes densely crowded, or elongate with distant flowers.
Calyx tubular, 5-angled and ribbed on the angles, unequally 5-toothed, ± dilated at the base in the fruiting stage and then ventricose, eventually splitting.
Leaves opposite and usually decussate, rarely 3-whorled or alternate, dentate, or often incised or variously lobed or partite, rarely entire.
Flowers small or medium-sized, sessile, each solitary in the axil of a narrow bract.
Stems 4-angled, erect ascending or procumbent, glabrous or hairy.
Annual or perennial herbs, small shrubs, woody climbers or trees.
Seeds oblong; endosperm lacking or scanty.
Life form
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) 0.1 - 0.5
Mature height (meter) 0.1 - 0.5
Root system tap-root
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light 7-9
Soil humidity -
Soil texture -
Soil acidity 2-8
Soil nutriment -
Hardiness (USDA) 10-11

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings or seedlings. Seeds needs stratification.
Mode cuttings seedlings
Germination duration (days) 14 - 90
Germination temperacture (C°) 18
Germination luminosity dark
Germination treatment stratification
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Verbena unspecified picture
Verbena unspecified picture

Distribution

Verbena world distribution map, present in Australia, China, New Zealand, Panama, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:330551-2
WFO ID wfo-4000040140
COL ID 99FT5
BDTFX ID 87383
INPN ID 198880
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Obletia Patya Glandularia Verbena

Lower taxons

Verbena x allenii Verbena barbata Verbena berteroi Verbena x bingenensis Verbena bipinnatifida Verbena x blanchardii Verbena canadensis Verbena canescens Verbena caniuensis Verbena carnea Verbena carolina Verbena chiricahensis Verbena x deamii Verbena delticola Verbena dissecta Verbena ehrenbergiana Verbena filicaulis Verbena flava Verbena glabrata Verbena gooddingii Verbena x goodmanii Verbena gracilis Verbena herteri Verbena x illicita Verbena incompta Verbena intermedia Verbena kuntzeana Verbena laciniata Verbena landbeckii Verbena lilacina Verbena lilloana Verbena lindbergii Verbena lindmanii Verbena lobata Verbena macdougalii Verbena maritima Verbena menthifolia Verbena neomexicana Verbena ovata Verbena paraguariensis Verbena paranensis Verbena parodii Verbena perennis Verbena x perriana Verbena x perturbata Verbena peruviana Verbena phlogiflora Verbena platensis Verbena plicata Verbena polyantha Verbena pumila Verbena rectiloba Verbena regnelliana Verbena reitzii Verbena sagittalis Verbena santiaguensis Verbena scabra Verbena scrobiculata Verbena selloi Verbena sessilis Verbena simplex Verbena stellarioides Verbena strigosa Verbena tampensis Verbena verecunda Verbena xutha Verbena amoena Verbena andalgalensis Verbena araucana Verbena aurantiaca Verbena elegans Verbena x engelmannii Verbena ephedroides Verbena gynobasis Verbena halei Verbena hispida Verbena humifusa Verbena macrosperma Verbena marrubioides Verbena megapotamica Verbena montevidensis Verbena multiglandulosa Verbena nana Verbena officinalis Verbena paulensis Verbena quadrangulata Verbena ribifolia Verbena x rydbergii Verbena sphaerocarpa Verbena subincana Verbena teucriifolia Verbena thymoides Verbena tomophylla Verbena tumidula Verbena valerianoides Verbena venturii Verbena californica Verbena cloverae Verbena cuneifolia Verbena demissa Verbena malmii Verbena pinetorum Verbena scabrella Verbena townsendii Verbena villifolia Verbena moranii Verbena calinfera Verbena falcata Verbena madrensis Verbena livermorensis Verbena ambrosiifolia Verbena hybrida Verbena concepcionis Verbena alata Verbena tweedieana Verbena aristigera Verbena bipinnatifida Verbena malpaisana Verbena turneri Verbena angustilobata Verbena brachyrhynchos Verbena guaibensis Verbena x clemensorum Verbena corymbosa Verbena jordanensis Verbena microphylla Verbena x ostenii Verbena racemosa Verbena rugosa Verbena sulphurea Verbena x uruguayensis Verbena lipozygioides Verbena mendocina Verbena x moechina Verbena moctezumae Verbena delicatula Verbena bajacalifornica Verbena balansae Verbena catharinae Verbena cheitmaniana Verbena dusenii Verbena goyazensis Verbena gracilescens Verbena grisea Verbena guaranitica Verbena hassleriana Verbena hatschbachii Verbena hirta Verbena porrigens Verbena radicata Verbena recta Verbena x stuprosa Verbena tecticaulis Verbena tenera Verbena litoralis Verbena supina Verbena stricta Verbena lasiostachys Verbena hastata Verbena bracteata Verbena rigida Verbena brasiliensis Verbena urticifolia Verbena bonariensis