Annual or perennial herbs. Leaves opposite to alternate, petiolate, simple. Capitula long-pedunculate, hemispherical, becoming reflexed in fruit, discoid or radiate, terminal, solitary or in loose cymes; involucral bracts numerous, 2-or 3-seriate, subequal, herbaceous; receptacle convex, pitted; paleae caducous. Ray florets uniseriate, female, fertile (rarely sterile), ligulate. Disc florets bisexual, usually 5-lobed; anthers obtuse at base, with a narrow acute apical appendage; style branches linear, with shortly hairy subulate terminal appendages. Achenes compressed, oblong to obovate, at least those of disc florets winged. Pappus of 2 bristle-like awns or absent.