Vigna unguiculata (L.) Walp.

Blackeyed pea (en)

Species

Angiosperms > Fabales > Fabaceae > Vigna

Characteristics

Herbaceous annual or perennial, erect, trailing or twining, cultivated or wild, chiefly in grassland, but also in open woodland and woodland edges. Root system shallow, adventitious or turbinate tubers, nodules common on all systems. Stems variable in thickness, patent, pubescent to aculeate, scabrid to glabrous, hairs colourless. Leaflets variously shaped, from obtusely trilobed to more or less triangular-rhomboid to elongate-hastate, texture variable from soft chartaceous to coriaceous, pubescence variable from strigose on both lamina surfaces to sparsely strigose on dorsal nerves only. Stipules typically spurred, i.e. appendaged below the point of attachment; variable from large, broadly ovate and most obvious, green, extremely strigose and ranging to hairy on ribs only or glabrous, to relatively small, lanceolate, brown and glabrous, all margins ciliate. Inflorescence axillary, a raceme varying from elongated with 4-7 flowers to contracted with 2 flowers, extrafloral nectaries alternating with flowers. Flowers blue, mauve, purple to pinkish mauve. Calyx campanulate, lobes longer or shorter than tube, variably strigose, the two upper lobes connate up to half their length, rugose and plicate with age. Standard known to snap closed from the time of anthesis to a few hours later, buff or yellowish on outer surface so that the closed flower appears to be yellow as reported by many collectors, oblate, upper pair of calluses narrow, perpendicular, more or less parallel or divergent. Keel straight, obtuse, slightly canoe-shaped, paler mauve to white, contains no pockets. Stamens 9, connate to form an open tube in the lower part, free filaments of two lengths, anthers entangled with dorsal style brush so that they distribute pollen when forced through the rigid keel opening as it is suppressed by visiting insects, vexillary stamen free, geniculate at base. Nectary collar-shaped, lobed, surrounding base of ovary. Ovary pilose to pubescent or puberulous with long stiff hairs along the suture. Style elongation extended horizontally or up to an angle of ca. 45° beyond the stigma, flattened, rounded or truncate, 0.2-0.8(-1.0) mm long, the whole appearing hammer-shaped. Legumes erect or pendant, black to yellow, varyingly pubescent to glabrous, from ca. 200-300 mm in cultivated forms (900 mm in cv. gr. sesquipedalis) and ca. 60-100 mm long in the wild forms, beaked, twisted spirally for seed dispersal in wild forms, non-dehiscent in cultivated forms. Seeds variable in colour from white, red, light to dark brown or black, often mottled, size variable from 6-9 mm in cultivated forms, much smaller in wild forms, hilum eccentric, aril ring-like.
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Annual or perennial erect trailing or climbing herb, 1–3 m. long.. Stems striate, glabrous or slightly hairy particularly near the petiole-bases, sometimes with minute stiff hook-like hairs.. Leaflets 3, ovate, rhomboid or lanceolate, the laterals oblique, 1·5–16·5 cm. long, 1–12·5 cm. wide, all entire or terminal leaflets subhastate to 3-lobed at the base and laterals lobed on the outer margin, shortly acuminate or subacute and mucronulate, acute to rounded at the base, glabrous or sparsely pubescent on both surfaces; petioles 1·5–13 cm. long; rhachis 0·6–2·5 cm. long; petiolules 2–5 mm. long; stipules submedifixed, constricted at the point of attachment, upper part lanceolate, 0·6–2 cm. long, spur narrower, 2–6 mm. long.. Inflorescences axillary, few-several-flowered; rhachis 0·5–4(–5) cm. long; peduncle 2–36 cm. long, usually minutely uncinulate pubescent towards the apex; pedicels 1–2(–4) mm. long; bracts deciduous, lanceolate, 3–5 mm. long; bracteoles ± persistent, spathulate, 3–5 mm. long.. Calyx glabrous; tube 3–5·5 mm. long; lobes triangular-lanceolate, 2·5–14 mm. long, acuminate, ciliolate; upper pair joined for about half their length or only at the base.. Standard white, greenish, yellow or lilac-purple, paler outside, round, 1·2–3·3 cm. long, 1–3·2(–4) cm. wide, rounded or emarginate, glabrous; wings blue to purple; keel usually white or pale, not twisted.. Pods erect or eventually hanging in cultivated forms, linear-cylindrical, 5·5–10(–90 in some cultivated races) cm. long, 3–11 mm. wide, glabrous or minutely verruculose.. Seeds white to dark red or black, often mottled with black or brown, oblong or reniform, longest dimension 3·5–5(–12) mm., shorter dimension 2–3·5(–6·5) mm., 2·2(–4·5) mm. thick; hilum one-third to half the longest dimension, eccentric; rim-aril slightly developed.
Coarse erect or clambering herbs or vines; stems glabrous, drying terete or strongly angled. Leaves pinnate trifoliolate; leaflets ovate or lanceolate, the lat-eral leaflets strongly oblique, apically acute, acuminate or obtuse, basally obtuse, truncate or cordate, mostly 5-11 cm long, 1.5-5 cm wide, drying slightly discol-orous, glabrous except at the very base; petiolules ca. 3 mm long, pubescent; petiole and the rachis glabrous, drying angled, the petiole mostly exceeding the terminal leaflet; stipels elliptical, rounded, glabrous, the lateral pair 2 mm long, the terminal pair shorter; stipules large, ovate, acuminate, nervate, 7-16 mm long above insertion, produced into 2 oblong auricles to 5 mm long below insertion. Inflorescence 7-30 cm long, the few flowers congested at the apex; peduncle glabrous, erect; bracteoles 4-7 mm long, exceeding the calyx tube, glabrous, nervate, cucullate; bracts 2-3 mm long, glabrous, nervate, cucullate; pedicels stout, 1 mm long. Flowers various colors; calyx tubular campanulate, glabrous, 6-8 mm long, the upper pair fused, deltoid, the lower teeth acuminate, ca. 2 mm long; standard 1.5-2 cm long. Legume linear, straight or curved, slightly com-pressed, mostly 10-40 cm long, ca. 1 cm wide; seeds of various colors. This species is distinct in its glabrous stems leaves and foliage, in its large stipules which are produced below the insertion, and in its large bracteoles which are longer than the bracts or the calyx tube. The pods occasionally grow to great length, over 1 m long, but are mostly much shorter.
Annual or perennial erect trailing or climbing herb, 0.5-3 m long. Stem striate, glabrous, scabrous, pubescent or hairy. Leaflets 3, 1.5-16.5 x 1-12.5 cm, ovate, rhombic or lanceolate to linear, the laterals oblique, all entire or terminal leaflets subhastate to deeply 3-lobed at the base and laterals lobed on the outer margin, shortly acuminate or subacute and mucronulate, acute to rounded at the base, glabrous to pubescent on both surfaces; petiole 1.5-13 cm long; rhachis 0.6-2.5 cm long; stipules submedifixed, constricted at the point of attachment, multinerved, upper part lanceolate, 6-20 mm long, spur narrower, 2-6 mm long. Peduncle 2-36 cm long. Flower pink to purple, rarely white (mainly within var. unguiculata cultivar-group Melanophthalmus), 15-33 mm long; pedicel 1-3 mm long, not expanding as the pod matures; bracteoles 3-6 mm long, spathulate or lanceolate, 1-3-nerved. Calyx glabrous or pubescent; tube 2-5.5 mm long; lobes subequal, ciliolate, the upper pair joined for about half their length to form a broad lobe bifid at the apex or only at the base. Standard with 2 parallel widely spaced appendages. Pod 8-10 x 2.5-5 mm (except within cultivated races), linear-cylindrical, glabrous, scabrous, or pubescent. Seed 4-6.5 x 2-4.5 mm (except within cultivated races); hilum one third to half the longest dimension, excentric; rim aril slightly developed.
A creeping bean type plant with straight firm pods. Both cover crop types (leafy) and grain types occur. There is a deep tap root and many branches occur from it in the surface of the soil. The root nodules are large and round. They can be 5 mm across. The leaves have 3 leaflets. The end leaflet can be 12-16 cm long. This leaflet is larger than the side leaflets. The side leaflets are assymetrical. The stipules at the base of the leaf are large and with spurs at their base. Flowers occur often in pairs on the end of long flowering shoots. This stalk can be 2 cm to 30 cm long. Only 2-4 flowers in each stalk produce pods. Flowers are white, yellow or blue. They are large and showy. The standard petal is 2-3 cm across. The pods are about 15 cm long. The seeds are white except for a dark scar.
Annual or perennial herbs, erect, trailing, or twining. Stems 1-3 m, subglabrous. Stipules lanceolate, ca. 1 cm, with a narrow spur below point of attachment; leaflets ovate-rhom­boid, 5-15 × 4-6 cm, lateral ones oblique, puberulent or gla­brous on both surfaces, base acute to rounded, apex acute. Ra­cemes axillary, with 2-6 flowers clustered at top of rachis. Calyx campanulate, 6-10 mm; teeth lanceolate. Standard yel­lowish white or violet, suborbicular, 1.2-3.3 × 1-3.2 cm, apex emarginate; wings blue to purple, subdeltoid; keel usually white or pale, not twisted. Legumes terete, 7.5-30(-90) × 0.6-1 cm. Seeds several, dark red or black, mottled with black or brown, oblong or reniform, 6-12 mm.
Annual or perennial, herbaceous climber, 0.2-2.0 m high/long; stems variable in thickness and indumentum. Leaflets variously shaped, obtusely trilobed, triangular-rhomboid to elongate-hastate. Stipules extended with spur beyond point of attachment, variably shaped, margins ciliate. Inflorescences axillary, elongated (4-7 flowers) or contracted racemes (2 flowers). Calyx campanulate, lobes ± as long as tube. Style prolongation horizontal or upturned by 45°, hammer-shaped. Petals: standard subrotund; keel straight, without pockets. Flowering time Sept.-Apr. Pod 200-300 mm in cultivated forms, non-dehiscent; 60-100 mm in wild forms, beaked, twisted spirally.
Leaflets 3, 1.5–16.5 × 1–12.5 cm, ovate, rhombic or lanceolate to linear, the laterals oblique, all entire or terminal leaflets subhastate to deeply 3-lobed at the base and laterals lobed on the outer margin, shortly acuminate or subacute and mucronulate, acute to rounded at the base, glabrous to pubescent on both surfaces; petiole 1.5–13 cm long; rhachis 0.6–2.5 cm long; stipules submedifixed, constricted at the point of attachment, multinerved, upper part lanceolate, 6–20 mm long, spur narrower, 2–6 mm long.
Flower pink to purple, rarely white (mainly within var. unguiculata cultivar-group Melanophthalmus), 15–33 mm long; pedicel 1–3 mm long, not expanding as the pod matures; bracteoles 3–6 mm long, spathulate or lanceolate, 1–3-nerved.
Calyx glabrous or pubescent; tube 2–5.5 mm long; lobes subequal, ciliolate, the upper pair joined for about half their length to form a broad lobe bifid at the apex or only at the base.
Seed 4–6.5 × 2–4.5 mm (except within cultivated races); hilum one third to half the longest dimension, excentric; rim aril slightly developed.
Pod 8–10 × 2.5–5 mm (except within cultivated races), linear-cylindrical, glabrous, scabrous, or pubescent.
Annual or perennial erect trailing or climbing herb, 0.5–3 m long.
Stem striate, glabrous, scabrous, pubescent or hairy.
Stems annual but often stout, subglabrous, twining
Standard with 2 parallel widely spaced appendages.
Flowers white and mauve-tinged or pink or yellow.
Peduncle 2–36 cm long.
Life form annual
Growth form herb
Growth support climber
Foliage retention deciduous
Sexuality hermaphrodite
Pollination entomogamy
Spread -
Mature width (meter) 0.51 - 0.52
Mature height (meter) 0.8 - 2.0
Root system adventitious-root tap-root
Rooting depth (meter) 0.85
Root diameter (meter) 0.7
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer
Photosynthetic pathway c3

Environment

It grows in tropical and subtropical climates. It grows from sea level to 1800 metres altitude in the tropics. In Ethiopia it grows between 500-2,000 m above sea level. Plants can stand high temperatures. Some kinds can tolerate drought. They are sensitive to cold and killed by frost. Plants germinate with a temperature between 11.5-15.5°C. The best growth is between 20-35°C. They can grow on a range of soils providing they are well drained. They are a short day plant. They do well in the semiarid tropics. It will not tolerate acid or alkaline soils. It grows in areas with an annual rainfall between 280-410 mm. It can grow in arid places. It can tolerate shade so is good for inter-cropping.
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On trees, etc.
Light 5-8
Soil humidity 3-6
Soil texture 1-6
Soil acidity 5-8
Soil nutriment -
Hardiness (USDA) 9-12

Usage

The young leaves, young pods and ripe seeds are all eaten. They can be steamed, boiled, stir-fried etc. The leaves can be dried and stored. The dried seeds are used in soups and stews. They are ground into flour or fermented. The seeds are also used for bean sprouts. Roasted seeds are used as a coffee substitute.
Uses animal food coffee substitute cover crop environmental use fodder food food additive forage gene source green manure leaf vegetable manure material medicinal
Edible flowers fruits leaves pods roots seeds tubers
Therapeutic use Colic (seed), Cough (seed), Diuretics (seed), Anthelmintics (seed), Antipyretics (seed), Asthma (seed), Astringents (seed), Bronchitis (seed), Graves ophthalmopathy (seed), Heart diseases (seed), Hemorrhage (seed), Leukorrhea (seed), Lithiasis (seed), Menstruation disturbances (seed), Nephrolithiasis (seed), General tonic for rejuvenation (seed), Splenomegaly (seed), Tuberculosis, lymph node (seed), Urolithiasis (seed), Urticaria (seed), Rheumatoid arthritis (seed), Adenopathy (unspecified), Anus (unspecified), Astringent (unspecified), Bilious (unspecified), Breath (unspecified), Cyanogenetic (unspecified), Diarrhea (unspecified), Diuretic (unspecified), Dysuria (unspecified), Fistula (unspecified), Leprosy (unspecified), Liver (unspecified), Nausea (unspecified), Neuralgia (unspecified), Nutritive (unspecified), Polyuria (unspecified), Prolapse (unspecified), Rinderpest (unspecified), Stomach (unspecified), Thirst (unspecified), Tumor (unspecified), Jaundice (unspecified), Measles (unspecified), Pneumonia (unspecified), Sore (unspecified), Tonic (unspecified), Burn (unspecified), Dysentery (unspecified), Kidney (unspecified), Pleurisy (unspecified), Smallpox (unspecified), Anorexia (unspecified), Anthelmintics (unspecified), Aphrodisiacs (unspecified), Appetite stimulants (unspecified), Asthenia (unspecified), Burns (unspecified), Colic (unspecified), Diuretics (unspecified), Galactogogues (unspecified), Helminthiasis (unspecified), Hepatomegaly (unspecified), Hyperhidrosis (unspecified), Lactation disorders (unspecified), Leukorrhea (unspecified), Liver diseases (unspecified), Lymphadenitis (unspecified), Menstruation disturbances (unspecified), Splenic diseases (unspecified), Urination disorders (unspecified), Wounds and injuries (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

It is grown from seeds. Seed collection is easy. Seeds remain viable for several years if carefully stored. A seeding rate of about 20 kg per ha is suitable and seed are sometimes broadcast then thinned. Cowpeas mostly inbreed giving pure lines.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment soaking
Minimum temperature (C°) -
Optimum temperature (C°) 25 - 35
Size -
Vigor -
Productivity -

Images

Leaf

Vigna unguiculata leaf picture by Joy Vivian Austin (cc-by-sa)
Vigna unguiculata leaf picture by Jamie Hebb (cc-by-sa)
Vigna unguiculata leaf picture by Chasing a Murderer (cc-by-sa)

Flower

Vigna unguiculata flower picture by Arshad Kazi (cc-by-sa)
Vigna unguiculata flower picture by pierre vernet (cc-by-sa)
Vigna unguiculata flower picture by Ha Nguyen (cc-by-sa)

Fruit

Vigna unguiculata fruit picture by Dimitry Novoklimov (cc-by-sa)
Vigna unguiculata fruit picture by Ajay Sahu (cc-by-sa)
Vigna unguiculata fruit picture by Dimitry Novoklimov (cc-by-sa)

Distribution

Vigna unguiculata world distribution map, present in Afghanistan, Angola, American Samoa, Australia, Burundi, Bonaire, Sint Eustatius and Saba, Bangladesh, Brazil, Botswana, China, Côte d'Ivoire, Cameroon, Comoros, Djibouti, Dominican Republic, Ethiopia, Fiji, Ghana, Guinea, Guinea-Bissau, Equatorial Guinea, Guatemala, French Guiana, Guyana, Haiti, Indonesia, India, Iran (Islamic Republic of), Iraq, Kenya, Cambodia, Lao People's Democratic Republic, Liberia, Sri Lanka, Madagascar, Mali, Myanmar, Mozambique, Mauritius, Malawi, Namibia, Niger, Nigeria, Nicaragua, Niue, Nepal, Pakistan, Panama, Peru, Philippines, Papua New Guinea, Réunion, Rwanda, Sudan, Singapore, Sierra Leone, El Salvador, Suriname, Seychelles, Chad, Togo, Thailand, Tanzania, United Republic of, Uganda, United States of America, Viet Nam, Yemen, South Africa, and Zambia

Identifiers

LSID urn:lsid:ipni.org:names:1127257-2
WFO ID wfo-0000189386
COL ID 7FSHX
BDTFX ID 83355
INPN ID 447217
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Dolichos unguiculatus Vigna catjang Vigna scabra Dolichos melanophtalmus Phaseolus unguiculatus Dolichos unguiculata Dolichos catiang Dolichos catjang Vigna catjang Phaseolus ellipticus Vigna sinensis subsp. sinensis Vigna unguiculata var. catjang Vigna sinensis subsp. cylindrica Vigna unguiculata var. catjang Vigna sinensis var. catjang Vigna unguiculata var. unguiculata Vigna catjiang Vigna sinensis var. spontea Vigna unguiculata

Lower taxons

Vigna unguiculata subsp. dekindtiana Vigna unguiculata subsp. sesquipedalis Vigna unguiculata subsp. stenophylla Vigna unguiculata subsp. tenuis Vigna unguiculata subsp. unguiculata Vigna unguiculata subsp. letouzeyi Vigna unguiculata subsp. burundiensis Vigna unguiculata subsp. pawekiae Vigna unguiculata subsp. pubescens Vigna unguiculata subsp. tenuis Vigna unguiculata subsp. aduensis Vigna unguiculata subsp. alba Vigna unguiculata subsp. baoulensis Vigna unguiculata var. ovata