Vigna Savi

Cowpea (en)

Genus

Angiosperms > Fabales > Fabaceae

Characteristics

Climbing, prostrate or erect herbs or subshrubs, rarely small shrubs, mostly from woody or tuberous rootstocks. Leaves pinnately, more rarely subdigitately, 3-foliolate or 1-foliolate; stipules truncate, bilobed or spurred at the base or sometimes quite peltate; stipels ± persistent, rarely absent. Inflorescences axillary or terminal, falsely racemose or flowers in dense 1-many-flowered subumbellate clusters or fasciculate; rhachis usually thickened and glandular at the point of insertion of the pedicels; bracts and bracteoles ± deciduous. Calyx 5-lobed, 2-lipped; lower lip 3-lobed,the middle lobe usually the longest; upper lip of 2 lobes completely or partly united. Corolla small or medium-sized, yellow, blue or purple; standard with inflexed auricles and 2–4 appendages or sometimes a single structure on the internal face, less often without appendages; keel truncate, obtuse or conspicuously beaked, sometimes the beak incurved through up to 360° (in some species the keel is twisted and untidy-looking and in others there is a distinct conical pocket on the left-hand petal). Vexillary stamen free; 5 shorter filaments (including the vexillary one) sometimes (in subgen. Haydonia) with a pair of joined glands below each anther; anthers uniform. Ovary 3–many-ovuled; style with tenuous lower part obsolete to quite long, filiform or flattened, upper part thickened and cartilaginous, straight or curved, upper portion barbate or hirsute on the internal face, sometimes produced beyond the stigma to form a short to long subulate or rarely flattened or capitate beak; stigma completely lateral, oblique or rarely ± terminal. Pods linear or linear-oblong, cylindrical or flattened, straight or ± curved, usually ± septate. Seeds mostly reniform or quadrate; hilum small or elongate; aril obsolete to well developed, usually eccentric, often 3-pronged.
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Vines or herbs, rarely shrubby, mostly with sturdy rootstocks, lacking unci-nate hairs. Leaves pinnate or subdigitate trifoliolate, rarely 1-foliate, mostly en-tire; stipels, nervate, mostly blunt; glabrate stipules sometimes produced below the insertion, sometimes bilobate. Inflorescences axillary or terminal, pseudor-acemes or subcapitate, the rachis contracted, the nodes glandular; bracts and bracteoles caducous; pedicels 1-2 per node, mostly shorter than the calyx. Flow-ers yellowish or white, sometimes with some purple or violet; calyx 2 lipped, the upper lip emarginate, the lowest tooth of the lower lip longest; standard orbicular, auriculate, sometimes appendaged on the dorsal face, the wings about equalling the standard and keel, the keel apically curved up to 1 spiral, often oblique; vexillary stamen free, the anthers uniform; style apically thickened and barbate on the inner face, caducous, sometimes with a curved beak. Legume linear to oblong, turgid or compressed, straight or curved, not septate; seeds reniform or quadrate, the hilum short or long, a well developed aril sometimes present. Vigna embraces about 150 species of tropical regions of both hemispheres. In Panama most species have flowers at least partly yellow, but in other countries some species have violet or purple flowers. A number of species of Vigna are important food plants for man, e.g. V. unguiculata, and species are sometimes cultivated for cattle fodder.
Corolla small or medium-sized, yellow, blue or purple inside (internal face of standard, external face of wings), greenish outside (external face of standard), all petals of subequal length; standard glabrous (except in V. heterophylla), emarginate, usually slightly wider than long and symmetrical, with inflexed auricles and appendages on the internal face, less often without appendages; appendages of the standard are based on a U-shaped pattern with one on each half of the standard, but the pattern is rarely complete; it can be reduced to the central part of the U with the appendages appearing parallel and very close together (central position), or sometimes joined and appearing V-or X-shaped (V.luteola or V. monophylla for example); it can be reduced to the lateral part of the U with the appendages appearing parallel but spaced apart (lateral position) (as in V. unguiculata); it can be reduced to the basal part of the U with the appendages appearing perpendicular to the standard axis (V. comosa); keel whitish except for the beak (if there is a beak), usually fused on the upper side, truncate, obtuse or conspicuously beaked, sometimes the beak incurved through up to 180° (V. radiata), sometimes with a distinct conical pocket on the left-hand petal (V. vexillata for example).
Twining or erect herbs, rarely subshrubs. Leaves pinnately 3-foliolate; stipules peltate or basally spurred, 2-lobed, cordate, or truncate. Racemes axillary or terminal, nodes of rachis often thickened and glandular. Bracts and bracteoles deciduous. Calyx 5-toothed, 2-lipped. Corolla yellow, blue, or purple; standard suborbicular, base appendaged; wings shorter than standard; keel subequal to wings, incurved, not beaked or produced into an incurved or spirally twisted beak. Stamens diadelphous; anthers uniform. Ovary sessile; style filiform, upper part thickened, bearded or hirsute lengthwise inside; stigma oblique. Legumes linear or linear-oblong, terete or flat. Seeds reniform or subquadrate; hilum short or elongate, with or without aril.
Inflorescence axillary, falsely racemose or flowers in dense 1–many-flowered subumbellate clusters or fasciculate; rhachis thickened and glandular at the point of insertion of the pedicels, flowers paired at each node; bracts and bracteoles deciduous, usually similar in shape and nervation; pedicel shorter than or as long as the calyx, extending or not as the pod matures.
Ovary 1–many-ovuled; style with tenuous lower part obsolete to quite long, filiform or flattened, upper part thickened and cartilaginous, straight or curved, upper portion barbate or hirsute on the internal face, produced beyond the stigma to form a short to long subulate beak (except in subgenus Haydonia); stigma completely lateral or oblique.
Leaves pinnately, more rarely subdigitately, 3-foliolate, 1-foliolate or simple; leaflets entire, venation usually reticulate, rarely with secondary nerves parallel (V. multinervis) or tertiary nerves parallel; stipules usually bilobed or spurred at the base, sometimes peltate, rarely truncate; stipels persistent, rarely absent.
Seeds mostly reniform or quadrate, thickness usually slightly less than width, usually cream-coloured, cream-coloured in combination with grey, mottled and speckled patterns, or black; hilum small or elongate; aril obsolete to well developed, usually excentric, often 3-pronged.
Pods linear or linear-oblong, usually terete, rarely flattened (V. macrorhyncha), with sutures not raised (except V. macrorhyncha), straight or curved, not septate (seeds are separated by a spongy tissue, not as woody as in Dysolobium and Pachyrhizus); style caducous.
Vexillary stamen free; 5 shorter filaments (including the vexillary one) sometimes with a pair of joined glands below each anther (in subgenus Haydonia); anthers uniform.
Climbing, twining, prostrate or erect herbs or subshrubs, rarely small shrubs, mostly from woody or tuberous rootstocks, without hooked hairs (as in Phaseolus).
Calyx 5-lobed, 2-lipped; lower lip 3-lobed, the middle lobe usually the longest; upper lip of 2 lobes completely or partly united.
Pollen triporate, exine coarsely reticulate (except in subgenus Haydonia).
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Hardiness (USDA) 4-10

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Images

Vigna unspecified picture
Vigna unspecified picture

Distribution

Vigna world distribution map, present in China, Panama, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:325971-2
WFO ID wfo-4000040261
COL ID 86RX
BDTFX ID 87389
INPN ID 445695
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Callicysthus Rudua Voandzeia Azukia Geolobus Phasellus Plectrotropis Scytalis Vigna

Lower taxons

Vigna antunesii Vigna marina Vigna angularis Vigna debanensis Vigna lanceolata Vigna haumaniana Vigna juncea Vigna juruana Vigna kirkii Vigna longifolia Vigna mendesii Vigna multinervis Vigna nervosa Vigna nigritia Vigna oblongifolia Vigna platyloba Vigna pygmaea Vigna radicans Vigna racemosa Vigna richardsiae Vigna subterranea Vigna trilobata Vigna umbellata Vigna vexillata Vigna wittei Vigna minima Vigna reflexo-pilosa Vigna kokii Vigna mudenia Vigna schlechteri Vigna tisserantiana Vigna microsperma Vigna dalzelliana Vigna bournaea Vigna halophila Vigna procera Vigna lasiocarpa Vigna ambacensis Vigna pubigera Vigna comosa Vigna friesiorum Vigna hosei Vigna jaegeri Vigna laurentii Vigna lobatifolia Vigna membranacea Vigna mildbraedii Vigna monophylla Vigna nuda Vigna ramanniana Vigna reticulata Vigna schimperi Vigna somaliensis Vigna triphylla Vigna angivensis Vigna bosseri Vigna hirtella Vigna clarkei Vigna hainiana Vigna khandalensis Vigna mukerjeanus Vigna subramanianus Vigna malayana Vigna tenuicaulis Vigna mungo Vigna verdcourtii Vigna aconitifolia Vigna bequaertii Vigna radiata Vigna monantha Vigna gracilicaulis Vigna gracilis Vigna longissima Vigna indica Vigna lonchophylla Vigna owahuensis Vigna khandalensis Vigna myrtifolia Vigna halophila Vigna trichocarpa Vigna konkanensis Vigna triodiophila Vigna pandeyana Vigna sathishiana Vigna yadavii Vigna sahyadriana Vigna polytyla Vigna fischeri Vigna heterophylla Vigna parkeri Vigna phoenix Vigna keraudrenii Vigna nepalensis Vigna pseudovenulosa Vigna exilis Vigna grandiflora Vigna bourneae Vigna nyangensis Vigna glabrescens Vigna dolomitica Vigna gazensis Vigna filicaulis Vigna stenophylla Vigna venulosa Vigna truxillensis Vigna unguiculata Vigna frutescens Vigna luteola