Herbs, perennial, terrestrial, not rhizomatous. Stems erect or ascending, simple or branched. Leaves few, along stem, opposite, very unequal in each pair; leaf blade sparsely puberulent, base strongly oblique, cuneate to cordate. Inflorescences lax, axillary or pseudoterminal, few-to many-flowered cymes; bracts apparently absent. Calyx actinomorphic, 5-sect from base or 5-lobed; segments equal, with many longitudinal secretory canals. Corolla white, reddish, or purple, zygomorphic, inside sparsely puberulent below abaxial lip or with lines of hairs; tube subtubular to funnelform-tubular, not swollen, longer than limb, 4-6 mm in diam.; limb 2-lipped; adaxial lip 2-lobed, slightly shorter than to subequal to abaxial lip; abaxial lip 3-lobed, lobes equal or central lobe longer, apex rounded. Stamens 4, adnate to corolla tube near base, included; anthers dorsifixed, coherent, thecae divergent, confluent at apex, dehiscing longitudinally; connective not projecting; staminode 1, adnate to adaxial side of corolla tube. Disc ringlike. Ovary ovoid to depressed globose, 2-loculed; placenta 1, axile, projecting inward, 2-cleft. Stigmas 2, equal, suborbicular, undivided, or 1, terminal, disclike, undivided. Capsule straight in relation to pedicel, subglobose, globose, or depressed globose, shorter than calyx, dehiscing loculicidally to base; valves 2, or dehiscing irregularly. Seeds unappendaged.