Stems compact, erect to ascending, with a few persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. Leaves 5--25 × 1.5--6 cm. Petiole reddish brown to dark purple when mature, not articulate above base, somewhat pliable and resistant to shattering. Blade lanceolate to ovate, usually 2-pinnate proximally, densely glandular, often somewhat viscid; most glandular hairs with thick stalks and distinctly bulbous tips; rachis with abundant glandular hairs and a few narrow scales. Pinnae ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 5--11 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. Pinnules dentate, often shallowly lobed; margins nonlustrous, thin, densely glandular, lacking cilia but with occasional 1--2-celled translucent projections. Vein tips slightly (if at all) enlarged, barely visible adaxially. Indusia of relatively broad segments; segments multiseriate for most of length, often divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia. Spores averaging 44--50 µm. 2 n = 152.
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