Woodwardia unigemmata (Makino) Nakai

Species

Pteridophytes > Polypodiales > Blechnaceae > Woodwardioideae > Woodwardia

Characteristics

Plants terrestrial. Stems stout, short-creeping to suberect, densely clothed at the apex with large, broadly to narrowly lanceolate, orange-brown scales; trophopods present. Fronds spreading, wintergreen and coriaceous, clustered near the stem apex, gemmiferous on the rhachis by the presence of one or two non-articulate scaly buds at or very near the frond apex, these gemmae usually developing 2-5 fronds and numerous roots while still attached. Stipes 32-100 cm long, stramineous to orange-brown, becoming darker toward the base, expanded at base, with two large adaxial vascular bundles and an arc of several smaller abaxial vascular bundles, densely scaly at the base and bearing scurfy remnants of multicellular glands, hairs, and scales distally, the scales sometimes leaving distinct scars after abscising, especially near the stipe base. Laminae 30-165 cm long, narrowly ovate, pinnate-pinnatifid, dark red to bronze or brown, glandular and scaly during emergence, becoming dark green and glabrous at maturity; pinnae in 11-16 pairs, the medial ones 18-61 cm and 2-10 cm, alternately arranged, deeply pinnatifid, lanceolate and acute at apex, basally ± symmetric about the costae, pinna lobes narrowly deltate, acute, separated from each other by a distinct sinus, coarsely serrulate along a strongly involute margin. Sori 1.5-3 mm long, linear-elongate to lunate, deeply immersed in the lamina, confined to the primary areoles of the costules; indusia cartilaginous with a thickened basal flange and a membranous conduplicate margin, vaulted, opening along a basal hinge like the lid of a box upon dehiscence of the sporangia. Spores 64, 30-35 by 2-23 µm (exclusive of the perispore), surrounded by a tightly appressed perispore presenting a reticulum of raised narrow ridges and rugulose expanses between the ridges, ornamented with aggregations of rods and cristae.
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Rhizome decumbent, dark brown, stout, up to 3 cm in diam., densely scaly; scales brown, lanceolate, 2.2-2.7 cm, membranous, entire, apex long acuminate. Stipes tufted, 30-100 cm, 5-8 mm in diam., base densely scaly; upper part of stipe and rachis with fewer brown, fibriform scales; lamina deeply bipinnatifid, elliptic or ovate-lanceolate, 30-100 × 20-60 cm, leathery; pinnae 7-15 pairs, subsessile or shortly stalked, broadly lanceolate; middle pinnae 15-35 × 4-12 cm, cut 3/4 way to costa, base rounded-truncate, unequal, apex caudate; lobes 14-20 pairs, close, oblique, oblong-lanceolate, sometimes falcate, lower pairs of lobes slightly shortened, middle lobes 1-6 × 0.8-1.2 cm, margin serrulate with sharp, long teeth; veins anastomosing with 2 or 3 rows of areoles along costae and costules, marginally free, simple or forked. Bulbils single or multiple, large, scaly, subterminal below rachis tip at base of upper pinna. Sori occupying costular areoles, oblong, sunken in rimmed depressions; indusia attached to outer vein, dark brown, thickly membranous. 2n = 68.
A fern.
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Growth form herb
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Mature width (meter) 1.5 - 2.5
Mature height (meter) 1.0 - 1.5
Root system rhizome
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Nitrogen fixer -
Photosynthetic pathway c3

Environment

Mixed deciduous, mixed deciduous-coniferous, and subtropical evergreen forests, nearly always occurring in humid, lightly shaded areas, often where the soil is shallow and rocky. Altitude 0-3000 m.
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It is a tropical plant.
Light 4-6
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Soil acidity 4-6
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Hardiness (USDA) 9-11

Usage

The starch in the rhizome is used for cakes and noodles.
Uses environmental use medicinal
Edible rhizomes roots
Therapeutic use -
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Cultivation

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Images

Woodwardia unigemmata unspecified picture

Distribution

Woodwardia unigemmata world distribution map, present in Bhutan, Botswana, China, India, Japan, Myanmar, Nepal, Pakistan, Philippines, Papua New Guinea, Taiwan, Province of China, and Viet Nam

Identifiers

LSID urn:lsid:ipni.org:names:17365530-1
WFO ID wfo-0001107452
COL ID 5C3CY
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Woodwardia yunnanensis Woodwardia himalaica Woodwardia latiloba Woodwardia maxima Woodwardia unigemmata Woodwardia radicans var. unigemmata