Agrimonia L.

Herbs, perennial, 3–20 dm, ?self-compatible?; rhizomatous, ?roots slender, fibrous, sometimes with thickened, ± fusiform or ± narrowly oblong tubers?. Stems 1–5, ascending to erect. Leaves persistent or basal deciduous and cauline persistent, cauline (see discussion); stipules persistent, adnate to petiole, ± 1/2-ovate, margins dentate or deeply incised, rarely entire; petiole present; blade ± narrowly obovate to ± narrowly elliptic, 1.5–22 cm, herbaceous, major leaflets 3–19, ?leaflets along rachis in ± opposite pairs, minor leaflet pairs between each major pair 0–5, number usually increasing distally along rachis?, major leaflet blades ovate, lanceolate, elliptic, rhombic, and/or obovate, minor leaflet blades ovate, elliptic, or obovate, margins flat, serrate to dentate, rarely incised, or in minor leaflet blades entire or few-toothed, eglandular hairs of two types: (1) soft and either straight or wavy (surfaces pilose, pubescent, or villous), and (2) stiff and straight (surfaces hirsute); glandular hairs of two types: (1) short-stipitate, and (2) sessile, both ± glistening (indumentum description applies to all plant surfaces in Agrimonia). Inflorescences terminal and often axillary, 9–120-flowered, simple or compound racemes (flexible); peduncles present; bracts present; bracteoles present. Pedicels present. Flowers 5–10 mm diam.; hypanthium ?stipitate, very short stipe adnate to hypanthium above bracteoles, reflexed at maturity, fruit abscising at base of stipe?, hemispheric to obconic, becoming indurate, 0.5–2 mm, ?sulcate, throat occluded by annular disc?, bristles in 2–5 circumferential rows from rim; sepals 5, spreading, ± ovate, 1–3 mm; petals 5, yellow, obovate to ± oval; stamens 5–15, shorter than petals; carpels 2, rarely more. Fruits achenes, 1 (1 ovary aborts), (including hypanthia) top-shaped, 1–6.6 mm; hypanthium persistent, enclosing achenes; sepals persistent, erect, connivent, ?bristles erect to reflexed, hooked?. x = 7.