Trees or shrubs, lacticiferous, the latex white or reddish. Leaves alternate, rarely opposite and decussate, or ternate, non-glandular, pinnately-nerved; mostly petiolate. Inflorescences cymose to aggregate dichasial and thyrsiform, axillary or terminal, sometimes extra-axillary, the secondary peduncles frequently fascicu-late in umbell-like form. Flowers small-to medium-sized; calyx 5-lobed, the lobes equal to conspicuously unequal, or rarely 4-lobed with the 2 outermost lobes con-nate and the 2 inner lobes free and included, without squamellae within; corolla salverform, tubular-salverform or tubular, the tube straight or inflated, ,terete or callose-angulate, the limb 5-lobed, sinistrorsely convolute; stamens 5, inserted midway or higher in the corolla tube, filamented, the anthers free and not agglu-tinated to the stigma, without an enlarged connective; gynoecium 2-carpellate, apocarpous, the ovary superior, the ovules numerous, rarely reduced to 2, with or without a low annular nectary, the style 1, the stigma ? clavate to subcapitate. Fruit apocarpous, of 2 follicles, + woody, mostly compressed, rarely cylindroid; seeds numerous, rarely reduced to 2, compressed, peltate, winged, the wing con-centric or eccentric.
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