Annual or perennial herbs, less often shrubs. Leaves opposite, scabrid or hispid. Capitula terminal, solitary or in lax cymes, or axillary and in congested corymbs, radiate; phyllaries 2–3-seriate, the outer herbaceous, the inner scarious; receptacle with conduplicate paleae. Ray florets with the ray 2-lobed at apex, with or without style, not producing mature achenes; disc florets tubular; anther bases obtuse or shortly sagittate, thecae and appendages often black; style branches gradually narrowing, pubescent at margins. Achenes obovoid, compressed, pubescent; pappus a laciniate cupule, often with a few setae.