Boerhavia L.

Spiderling (en), Punarnava (fr)

Genus

Angiosperms > Caryophyllales > Nyctaginaceae

Characteristics

Annual or perennial herbs, sometimes suffrutescent or subscandent; glabrous or pubescent. Leaves opposite or subopposite, those of a pair often unequal, large leaves often alternating; petiolate; blade entire, sinuate or undulate, raphides prominent, linear, base sometimes inequilateral. Inflorescences varying from a terminal or axillary panicle to paniculate capitula, or flowers arranged in racemes, unequally pedicellate umbels or cymes, flowers rarely solitary; branches often subtended by a small hyaline bract; pedicellate or subsessile; bracteoles 1-3, minute, hyaline. Flowers bisexual, small; perianth corolla-like, campanulate, subrotate or funnelform, lower part constricted above ovary, persistent, upper part (limb) shallowly 5-lobed, flaring, deciduous; stamens 1-3(-5), exserted or included, filaments free or connate below, pollen forate, 2(-3)-nucleate; ovary shortly stipitate, ovule anacampylotropous, style filiform, stigma 1, capitate, subexserted to exserted. Anthocarps shortly stipitate, clavate, fusiform or subellipsoid, 3-5(-10)-angled,-ribbed or-winged, green, with stipitate, viscous glands on ribs (eglandular in B. erecta, not in the Guianas), and streaks of white, linear superficial raphides in furrows between ribs; seed with thin testa adherent to pericarp, embryo with unequal cotyledons curved inward, enclosing scant, mealy endosperm, radicle elongate, descending.
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Herbs, annual or perennial, sometimes suffrutescent at base, slender, often glandular, glabrous, or pubescent, from slender and soft or stout, ± woody, and ropelike or fusiform taproot. Stems procumbent, decumbent, ascending, or erect, unarmed, with or without glutinous bands on internodes. Leaves petiolate, pairs unequal in size in each pair; blade thin or thick and slightly fleshy, base symmetric to asymmetric. Inflorescences terminal and axillary, pedunculate or not clearly pedunculate because of repeated branching from distal axils, diffuse, and then usually widely cymose, paniculate, or thyrsiform, terminal portions cymose, racemose, spicate, subumbellate, umbellate, subcapitate, or capitate, rarely borne singly; bracts ± persistent and not accrescent, or deciduous, 1-3 beneath each flower, distinct, lanceolate, minute, thin, translucent. Flowers bisexual, chasmogamous; perianth radially symmetric or slightly bilaterally symmetric, campanulate or widely funnelform, constricted beyond ovaries, tube abruptly expanded to (4-)5-lobed limb; stamens 2-8, included or exserted; styles at or extending beyond anthers; stigmas peltate. Fruits fusiform, clavate, oblong-clavate, obovoid, or obpyramidal, stiffly coriaceous; ribs (3-)5, rounded, angular, or winglike, smooth, glabrous or glandular-pubescent; sulci smooth or rugose, epidermal surface smooth, papillate, or minutely pubescent.
Annual herbs, erect, ascending or creeping, puberulous-glabrescent, with sessile or stalked, club-shaped glands or hairs; stem base and root often woody. Stems often red tinged and swollen (when dry constricted) at the nodes. Leaves opposite, subequal in each pair, beneath paler, the epidermis with minute irregular cystolith-like sculpture, and sometimes with embedded reddish glands. Inflorescences axillary, in the axil of the smallest leaf of each pair, or (B. erecta) by reduction of leaves into bracts each stem forming one large thyrsoid inflorescence appearing terminal at the extremities, subumbels of 2-10 small flowers. Bracts (basal) and bracteoles (apical) small, acute, fimbriate, caducous. Pedicels jointed with the flower, mostly very short. Flowers bisexual. Perianth tubular-campanulate, with a distinct constriction mostly halfway; lower part (later becoming the coriaceous anthocarp) obconical, 5-or 10-ribbed, upper caducous part 5-lobed, plicate in bud, white or pink. Stamens 1-4, exserted. Ovary (sub)sessile, smooth; style as long as the perianth; stigma capitate. Anthocarp closed at apex, 5-10-ribbed, glabrous or set with glands, swelling and slimy in water. Seed with longitudinally folded embryo; cotyledons with recurved margin and surrounding the mealy endosperm.
Annual or perennial herbs, erect, diffuse or decumbent, sometimes with woody base; roots commonly thick, fleshy or woody; stems usually branched, terete, glabrous or hairy; hairs glandular, often glandular-viscid; raphides commonly present in all structures. Leaves opposite, those at each node often of unequal size, sinuate or entire, often undulate, often fleshy, usually grading to bracts above. Inflorescences usually grading along branches from simple to compound; basic inflorescence unit a solitary flower, a glomerule, or an umbel; bracteoles minute, membranous. Flowers small, bisexual. Perianth tubular, the upper part caducous, 5-lobed, campanulate or trumpet-shaped, white, pink, mauve or crimson-purple. Stamens 1–4, rarely to 6; anthers suborbicular, included or shortly exserted. Style slender, often elongate; stigma capitate. Fruit indehiscent, conspicuously 3–5-sulcate, with blunt, acute or winged ribs, often densely glandular-viscid; ribs usually mucous when wet. Seed 1, erect. See also Du Puy & Telford (1993: 104), Green (1994: 75).
Herbs annual or perennial, erect, ascending, or procumbent. Stems sometimes glandular. Leaves opposite, often unequal, petiolate, margin entire or sinuate. Inflorescences usually terminal, sometimes apparently axillary, of small umbels, often irregular, rarely 1-flowered, grouped into cymose panicles. Flowers bisexual, small; bracteoles minute, often deciduous. Perianth constricted beyond the ovary, limb campanulate, apex 5-lobed, truncate, or plicate, caducous, Stamens 1-5, included or shortly exserted; filaments connate at base. Ovary oblique, stipitate. Stigma peltate or capitate. Fruit obovoid, turbinate, clavate, or terete, small, 5-ribbed or deeply 5-angled, often rough, with sessile, sticky glands. Seed: embryo curved, cotyledons thin, broad, enclosing the thin endosperm.
Annual or perennial herbs. Leaves opposite or subopposite, petiolate, frequently unequal at the nodes, the blade undulate or sinuate, conspicuously raphidulous. Inflorescence terminal, paniculiform to racemiform, many-flowered. Flowers small, perfect, subtended by 1-3 minute bracteoles, the perianth tube persistent, more or less herbaceous, frequently glandular, the perianth limb deciduous, petalaceous, obscurely 5-lobed; stamens 1-5, included or exserted; ovary shortly stipitate, the stigma capitate, more or less exserted. Anthocarp obpyramidal to obovoid or subellipsoid, glabrous to glandular, 3-to 5-angulate.
Life form annual
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
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Root system tap-root
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Flower color
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Nitrogen fixer -
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Environment

Distinctly heliophilous weeds of beaches and ruderal places preferring a slightly seasonal climate, indifferent to soil, up to c. 1000 m.
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Hardiness (USDA) 5-12

Usage

The fleshy roots of some species are reported to be eaten by the Aborigines.
Uses -
Edible -
Therapeutic use -
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Animal toxicity -

Cultivation

Mode -
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Images

Boerhavia unspecified picture
Boerhavia unspecified picture

Distribution

Boerhavia world distribution map, present in Australia, China, Panama, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:326857-2
WFO ID wfo-4000004873
COL ID 3B2R
BDTFX ID 86007
INPN ID 445856
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Boerhavia

Lower taxons

Boerhavia hualienensis Boerhavia acutifolia Boerhavia alamasona Boerhavia alata Boerhavia albiflora Boerhavia angustifolia Boerhavia anisophylla Boerhavia gracillima Boerhavia graminicola Boerhavia grandiflora Boerhavia greenwayi Boerhavia heimerlii Boerhavia helenae Boerhavia herbstii Boerhavia hereroensis Boerhavia heronensis Boerhavia hiranensis Boerhavia hirsuta Boerhavia lantsangensis Boerhavia linearifolia Boerhavia mista Boerhavia montana Boerhavia octandra Boerhavia organensis Boerhavia paludosa Boerhavia parviflora Boerhavia patula Boerhavia pedunculosa Boerhavia periplocifolia Boerhavia pilosa Boerhavia ramosissima Boerhavia raynalii Boerhavia rosei Boerhavia scandens Boerhavia sinuata Boerhavia stellata Boerhavia stenocarpa Boerhavia tuberosa Boerhavia virgata Boerhavia weberbaueri Boerhavia arabica Boerhavia boissieri Boerhavia brandegeei Boerhavia burbidgeana Boerhavia capitata Boerhavia chinensis Boerhavia chrysantha Boerhavia ciliata Boerhavia cordobensis Boerhavia coulteri Boerhavia crassifolia Boerhavia crispa Boerhavia diandra Boerhavia dichotoma Boerhavia discolor Boerhavia elegans Boerhavia fallacissima Boerhavia fistulosa Boerhavia gardneri Boerhavia glabrata Boerhavia glandulosa Boerhavia litoralis Boerhavia maculata Boerhavia megaptera Boerhavia mutabilis Boerhavia pentandra Boerhavia plumbaginea Boerhavia procumbens Boerhavia pulchella Boerhavia punarnava Boerhavia purpurascens Boerhavia reniformis Boerhavia repens Boerhavia repleta Boerhavia rufopilosa Boerhavia schinzii Boerhavia schomburgkiana Boerhavia simonyi Boerhavia sonorae Boerhavia spicata Boerhavia squarrosa Boerhavia tarapacana Boerhavia traubae Boerhavia triquetra Boerhavia tsarisbergensis Boerhavia verbenacea Boerhavia wrightii Boerhavia hualienense Boerhavia australis Boerhavia bracteosa Boerhavia crispifolia Boerhavia deserticola Boerhavia plicata Boerhavia xantii Boerhavia africana Boerhavia ambigua Boerhavia decipiens Boerhavia hitchcockii Boerhavia lateriflora Boerhavia libyca Boerhavia tetrandra Boerhavia transvaalensis Boerhavia coccinea Boerhavia diffusa Boerhavia erecta