Stamens (4)5, slightly perigynous, inserted between and united with the disk lobes, with filaments dorsally compressed, slightly dilated near the base; anthers oblong-sagittate, versatile.
Ovary narrowly ovoid-pyramidal, 5–10-angled, completely or incompletely 5(6–7)-locular, with 8 horizontal ovules in 2 rows; style thick, simple; stigma shallowly 5(6–7)-lobed.
Leaves alternate, entire or dentate to spinose-dentate, petiolate; stipules minute, caducous.
Flowers in condensed pedunculate cymes or false umbels or fascicles, sometimes cauliflorous.
Fruit drupaceous, woody, 5(6–7)-locular or eventually 1-locular, each loculus 8-seeded.
Disk (4)5-lobed, each lobe bearing 3–4(–6) stiff filaments (staminodes?).
Seeds obovoid-oblong, angular, with black testa; embryo as long as seed.
Petals (4)5, free, imbricate, spreading, deciduous.
Sepals (4)5, free, imbricate, deciduous.
Trees or shrubs, glabrous.