Trees, shrubs, shrublets or woody climbers, without tendrils, glabrous or with simple hairs, unarmed or with axillary shoots terminating in a spine, sometimes with rubber-like latex (gutta) in various parts appearing as elastic threads when a leaf is broken. Leaves simple, alternate or spiral to subopposite or opposite, sometimes fasciculate on short shoots, entire or with crenate or denticulate to spinose margins, penninerved; stipules small, simple or laciniate, sometimes united by a transverse ridge, usually deciduous, or absent. Flowers bisexual, polygamous or unisexual, monoecious or dioecious, actinomorphic, often fragrant, in axillary and/or terminal dichasial or monochasial cymes or panicles or thyrses, sometimes with accessory branches (additional to the normal one in the bract axil), or fasciculate or solitary, usually bracteate; pedicels often articulated. Sepals (3–)4–5(–6), imbricate or rarely valvate in bud, free or united at the base, persistent. Petals (3–)4–5(–6), free or rarely united at the base, imbricate or rarely valvate in bud, usually persistent, sometimes with ventral grooves or hollows or appendages. Stamens (2–)3–5(–10), antisepalous, free or more rarely with filaments partly united to form a tube, inserted outside or on or inside the disc (when present); anthers usually short, (1–)2-thecous, extrorse or introrse, basifixed or dorsifixed or versatile, sometimes deciduous, dehiscing by longitudinal or oblique or horizontal slits; pollen simple or more rarely in tetrads or polyads. Disc nectariferous, annular, entire or angular or crenulate or lobed or covered with fleshy processes, concave to convex, rarely wholly or partly incorporated in an androgynophore or formed of discontinuous pockets, fleshy or membranous, very rarely absent. Ovary free or partly or wholly immersed in the disc, sessile or on a short androgynophore, syncarpous, completely or very rarely incompletely 2–5-locular, or rarely 1-locular by abortion, with 1–many erect or rarely pendulous ovules in 2 rows or rarely superimposed in each locule; styles as many as the locules, free or ± united, or absent; stigmas various, free or ± united. Fruit capsular, loculicidal, or of divergent ± dorsally flattened dehiscent mericarps or baccate or drupaceous or dry, indehiscent and sometimes winged. Seeds with a fleshy or submembranous brightly coloured aril, or winged with the funicle free from the wing or united to its base, or neither arillate nor winged (usually in indehiscent fruits), with or without endosperm; embryo erect, with cotyledons flat or fleshy, rarely united
Trees, erect or scandent shrubs, lianas, evergreen or deciduous; stems sometimes producing rootlets (Euonymus spp.), sometimes thorny (Maytenus spp.), rarely with buttressed trunks. Leaves simple, alternate or opposite, petiolate; leaf blade laminar, venation pinnate, secondary veins reticulate, margins entire, crenate, serrate, or dentate; stipules small and caducous, or estipulate. Inflorescences axillary or terminal, few to many flowered, cymose, thyrsoid, racemose, fasciculate, or flowers solitary, rarely paniculate. Flowers actinomorphic, bisexual or unisexual; perianth (3 or)4-or 5-merous; sepals and petals free. Disk intrastaminal, stamens on disk, or extrastaminal, annular, margins upturned, pulvinate, or cupular, conspicuous, rarely obscure (Microtropis spp.), entire, lobed, or angular. Stamens 3-5, alternate with petals; anther (1 or)2-celled, basifixed to dorsifixed, dehiscing longitudinally or obliquely, introrse, extrorse, or latrorse. Ovaries 3-5, superior to half-inferior, often partially immersed in disk, completely or incompletely 2-5-locular, placentation axile, ovules erect, axile, or pendulous, ovules (1 or)2(or 3 or more) per locule; style terminal, simple, short to absent; stigma simple or lobed. Fruit a loculicidally dehiscent capsule, schizocarp of 2-5 indehiscent mericarps, drupe, berry, or samara with a single surrounding wing, rarely an indehiscent capsule or nut with lateral style, pericarp bony, leathery, chartaceous, or fleshy, capsules smooth, angular, deeply lobed, transversely flattened and lobed to base, or connate, rarely prickly, laterally winged. Seeds 1-12, smooth or occasionally furrowed, albuminous or exalbuminous, sometimes winged, wing membranous, basal, exarillate or aril basal to completely enveloping seed, aril membranous, fleshy, rarely mucilaginous; cotyledons flat, foliaceous or thick, connate, germination epigeous. 2n = 8, 12, 14.
Herbs, shrubs, trees, or vines, annual or perennial, deciduous or evergreen, synoecious, dioecious, or polygamomonoecious. Leaves alternate, subopposite, opposite, whorled, or fascicled, simple; stipules absent or present; petiole present or absent; blade margins serrate, dentate, spiny, or entire; venation pinnate, palmate, or 1-veined, sometimes obscure. Inflorescences unisexual or bisexual, terminal or axillary, cymes, racemes, panicles, thyrses, or fascicles, or flowers solitary. Flowers bisexual or unisexual, radially symmetric or weakly asymmetric; perianth and androecium hypogynous or perigynous; hypanthium free, completely adnate to ovary, or absent; sepals (3–)4–5[–7], distinct or connate proximally; petals 0 or (3–)4–5[–7], distinct; nectary present, rudimentary, or absent; stamens 3–5[–10], distinct, free or adnate to nectary; anthers dehiscing by longitudinal slits; <staminodes 0 or [4–]5[–7]>; pistil 1, 1–5-carpellate, ovary superior <often embedded in nectary> to 1/2 inferior, 1–5-locular, placentation axile or parietal; ovules 1–2[–4] or 100–2000+ per locule, anatropous; styles 0, 1, or 3, connate proximally; stigmas 2–5. Fruits capsules, dehiscence loculicidal, drupes, or nutlike (small, hard-walled, indehiscent, 1-locular, and 1-seeded) [berries or samaras]. Seeds 1, 2, 40–70, or 100–2000+ per locule, <often winged or covered by brightly colored pulpy aril>.
Ovary free or partly or wholly immersed in the disk, sessile or on a short androgynophore, syncarpous, completely or very rarely incompletely 2–5-locular, or rarely 1-locular by abortion, with 1-? erect or rarely pendulous ovules in 2 rows or rarely superimposed in each loculus; styles as many as the loculi, free or ± united, or absent; stigmas various, free or ± united
Stamens (2)3–5(6–10), antisepalous, free or more rarely with filaments partly united to form a tube, inserted outside or on or inside the disk; anthers usually short, (1)2-thecous, extrorse or introrse, basifixed or dorsifixed or versatile, sometimes deciduous, dehiscing by longitudinal or oblique or horizontal slits; pollen simple or more rarely in tetrads or polyads
Seeds with a fleshy or submembranous brightly coloured aril, or winged with the funicle free from the wing (Tab. 80 figs 6–7) or united to its base (Tab. 85 fig. A8) or neither arillate nor winged (usually in indehiscent fruits), with or without endosperm; embryo erect, with cotyledons flat or fleshy, rarely united
Flowers bisexual or polygamous or unisexual, monoecious or dioecious, actinomorphic, often fragrant, in axillary and/ or terminal dichasial or monochasial cymes or panicles or thyrses, sometimes with accessory branches, or fasciculate or solitary, usually bracteate; pedicels often articulated
Leaves simple, alternate or spiral to subopposite or opposite, sometimes fasciculate on short shoots, entire or with crenate or denticulate to spinose margins, penninerved; stipules small, simple or laciniate, sometimes united by a transverse ridge, usually deciduous, or absent
Trees, shrubs, shrublets or woody climbers, without tendrils, glabrous or with simple hairs, unarmed or with axillary shoots terminating in a spine, sometimes with rubber-like latex (gutta) in various parts appearing as elastic threads when a leaf is broken
Disk nectariferous, annular, entire or angular or crenulate or lobed or covered with fleshy processes, concave to convex, rarely wholly or partly forming an androgynophore or discontinuous pockets, fleshy or membranous, very rarely absent
Petals (3)4–5(6), free or rarely united at the base, imbricate or rarely valvate in bud, usually persistent, sometimes with ventral grooves or hollows or appendages
Fruit capsular, loculicidal, or of divergent ± dorsiventrally flattened dehiscent mericarps or baccate or drupaceous or dry, indehiscent and sometimes winged
Fruit a loculicidal capsule, an indehiscent fleshy or hard drupe, a berry, a 3-lobed capsule or a capsule with 3 nearly separate mericarps
Sepals (3)4–5(6), imbricate or rarely valvate in bud, free or united at the base, persistent
Small trees, shrubs or woody climbers without tendrils, sometimes spiny
Seed sometimes arillate, sometimes winged, sometimes neither
Carpels 3–5 united, free from or half-embedded in the disk
Leaves simple, alternate or opposite
Stamens 3–5, alternate with petals
Stipules inconspicuous or absent
Flowers bisexual or unisexual
Ovules 2–12 in each carpel
Sepals 4–5, usually free
Petals 4–5, free