Euonymus L.

Spindletree (en), Fusains véritables (fr), Fusain (fr)

Genus

Angiosperms > Celastrales > Celastraceae

Characteristics

Usually shrubs or small trees, rarely of moderate size (25 m), erect, rarely scandent, glabrous, evergreen or deciduous. Leaves opposite, usually decussate, very rarely spiral, alternate or verticillate (extra-Mal. spp.), sometimes with dark dots underneath. Stipules lanceolate, caducous. Inflorescences axillary, cymose, rarely flowers in fascicles (E. javanicus); pedicels articulated. Flowers bisexual, 5-or 4-merous. Calyx deeply lobed, imbricate, entire, erose, or minute-denticulate. Petals imbricate, spreading or reflexed, entire, erose, short-ciliate, or long-flmbriate, smooth or finely areolate (strong lens!) on the inner surface. Disk distinct, fleshy or thin, flat, 5-or 4-angular, or 5-or 4-lobed, or rounded, very rarely membranous cupular (in the African E. congolensis WILCZEK), smooth, or covered with fleshy papilla-like or subulate processes. Stamens inserted on the disk, at the margin or halfway; anthers deltoid, ± depressed-oblong, or broad-ovoid, obtuse or short-apiculate, slightly free at the base, dehiscent on the top, lateral, or introrse, either 2-celled or 1-celled (extra-Mal.); filaments obscure or distinct. Free part of the pistil terete, or slightly 4-5-angular towards the base, gradually narrowed to the apex; the basal part sometimes covered with papillose or subulate processes. Stigma obscure, obtuse, or discoid; ovary partly or ± wholly immersed in the disk; if the disk is angular the angles are episepalous. Ovules mostly 2 in each cell (rarely 3-12 in each cell in a few extra-Mal. spp.), attached to the inner angle near the base, or pendulous. Fruit a loculicidal capsule, often coloured, when dehiscing the valves splitting and leaving no central axis (except in the extra-Mal. sp. E. latifolia (L.) MILL.), usually (3-)4-5-angular or-lobed, sometimes globose, rarely fusiform, smooth or echinate; apex obtuse, acute, ± truncate, or concave. Seeds (usually black) with (mostly orange) aril at the base, or enveloped by it.
More
Shrubs, trees, or vines, <climbing by adventitious roots>. Branchlets terete or 4-angled. Leaves deciduous or persistent, opposite; stipules present; petiole present; blade margins entire or toothed; venation pinnate. Inflorescences terminal or axillary, cymes. Flowers bisexual, radially symmetric; perianth and androecium hypogynous; hypanthium absent; sepals 4–5, distinct; petals 4–5, white, green, yellow, red, or purple; nectary intrastaminal, annular, fleshy; stamens 4–5, adnate to nectary margin; staminodes 0; pistil 4–5-carpellate; ovary superior, 4–5-locular, placentation axile; style 1; stigma 1; ovules 2 per locule. Fruits capsules, (1–)2–5-locular, globose, subglobose, or obovoid, unlobed or 2–5-lobed, apex not beaked. Seeds 2 per locule, ellipsoid, ovoid, or subglobose, not winged; aril yellow, orange, or red, completely surrounding seed. x = 16.
Shrubs, sometimes small trees, ascending or clambering, evergreen or deciduous, glabrous, rarely pubescent. Leaves opposite, rarely also alternate or whorled, entire, serrulate, or crenate, stipulate. Inflorescences axillary, occasionally terminal, cymose. Flowers bisexual, 4(or 5)-merous; petals light yellow to dark purple. Disk fleshy, annular, 4-or 5-lobed, intrastaminal or stamens on disk; anthers longitudinally or obliquely dehiscent, introrse. Ovary 4-or 5-locular; ovules erect to pendulous, 2(-12) per locule. Capsule globose, rugose, prickly, laterally winged or deeply lobed, occasionally only 1-3 lobes developing, loculicidally dehiscent. Seeds 1 to several, typically 2 developing, ellipsoid; aril basal to enveloping seed.
Fls perfect, 4–6-merous, small, solitary or cymose in the axils; pet widely spreading; stamens very short, inserted at the margin of the broad disk, in which the 3–5-locular ovary is largely immersed; ovules 2–6 per locule, ± ascending; stigma sessile, 3–5-lobed; fr colored, capsular, often 3–5-lobed, with 1–6 seeds per locule, each seed covered by a red or orange aril; shrubs or small trees, prostrate to erect, sometimes climbing by adventitious roots; twigs often 4-angled, sometimes winged; lvs opposite, deciduous in our native spp. 200+, cosmop. Several cult. spp. that only occasionally escape are included in the key, but not numbered or described.
Small trees or shrubs, bisexual. Leaves opposite, rarely (not in Australia) otherwise; stipules present, caducous. Flowers in axillary cymes, rarely (not in Australia) in fascicles; pedicels articulate. Sepals 4 or 5, imbricate. Petals 4 or 5, imbricate. Disc present, angular, lobed or rounded. Stamens as many as petals and alternate with them, inserted on disc margin or inside; anthers latrorse or introrse. Ovary partly or wholly immersed in disc, 3-5-locular with 2-8 ovules per locule in two rows, attached to inner angle near base or pendulous; style 1; stigma 1. Fruit a capsule; columella present or absent. Seeds 1-several per locule, arillate.
Evergreen or deciduous, shrubs or small trees, occasionally creeping or ascending by rootlets. Lvs usually opposite, petiolate, glabrous. Fls 4-5-merous, in axillary cymes. Fr. a 4-5-celled capsule, usually lobed or winged. Seeds 0-2 per cell, usually covered by a brightly coloured fleshy aril.
Life form -
Growth form shrub
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 25.0
Root system adventitious-root creeping-root
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

In Malaysia in primary and secondary forests from the lowland up to 3200 m.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 4-10

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) 220 - 365
Germination temperacture (C°) 10 - 18
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Euonymus unspecified picture

Distribution

Euonymus world distribution map, present in Australia, China, Congo, Japan, Madagascar, Malaysia, New Zealand, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:331601-2
WFO ID wfo-4000014199
COL ID 4G5H
BDTFX ID 86415
INPN ID 446552
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Euonymus Masakia Melanocarya Quadripterygium Kalonymus Vyenomus

Lower taxons

Euonymus aculeatus Euonymus aculeolus Euonymus acuminifolius Euonymus balansae Euonymus bullatus Euonymus carnosus Euonymus centidens Euonymus crenulatus Euonymus elegantissimus Euonymus ficoides Euonymus frigidus Euonymus impressus Euonymus indicus Euonymus lichiangensis Euonymus macropterus Euonymus mexicanus Euonymus myrianthus Euonymus nanoides Euonymus nanus Euonymus occidentalis Euonymus pleurostylioides Euonymus potingensis Euonymus recurvans Euonymus rehderianus Euonymus sachalinensis Euonymus schensianus Euonymus semenovii Euonymus vaganoides Euonymus vagans Euonymus velutinus Euonymus venosus Euonymus verrucocarpus Euonymus viburnoides Euonymus barberi Euonymus leishanensis Euonymus wilsonii Euonymus acanthocarpus Euonymus acanthoxanthus Euonymus angulatus Euonymus attenuatus Euonymus australianus Euonymus benthamii Euonymus bockii Euonymus castaneifolius Euonymus cochinchinensis Euonymus contractus Euonymus corymbosus Euonymus costaricensis Euonymus darrisii Euonymus dichotomus Euonymus dielsianus Euonymus distichus Euonymus dolichopus Euonymus eberhardtii Euonymus echinatus Euonymus elaeodendroides Euonymus europaeus Euonymus gibber Euonymus giraldii Euonymus gracillimus Euonymus grandiflorus Euonymus hainanensis Euonymus hamiltonianus Euonymus huangii Euonymus jinyangensis Euonymus kachinensis Euonymus kanyakumariensis Euonymus kengmaensis Euonymus laxiflorus Euonymus lucidus Euonymus lutchuensis Euonymus maackii Euonymus macrocarpus Euonymus melananthus Euonymus mengtseanus Euonymus microcarpus Euonymus moluccensis Euonymus nitidus Euonymus obovatus Euonymus oxyphyllus Euonymus parasimilis Euonymus percoriaceus Euonymus petelotii Euonymus pittosporoides Euonymus sanguineus Euonymus serratifolius Euonymus subcordatus Euonymus subsulcatus Euonymus szechuanensis Euonymus tenuiserratus Euonymus ternifolius Euonymus theacola Euonymus tibeticus Euonymus tingens Euonymus walkeri Euonymus wrayi Euonymus wui Euonymus yunnanensis Euonymus phellomanus Euonymus baekdusanensis Euonymus chuii Euonymus clivicola Euonymus huae Euonymus phellomana Euonymus spraguei Euonymus tsoi Euonymus verrucosoides Euonymus glaber Euonymus tashiroi Euonymus theifolius Euonymus glandulosus Euonymus kweichowensis Euonymus enantiophyllus Euonymus maximowiczianus Euonymus lanceolatus Euonymus pseudovagans Euonymus yakushimensis Euonymus actinocarpus Euonymus cornutus Euonymus euscaphis Euonymus hupehensis Euonymus leiophloeus Euonymus tonkinensis Euonymus chiapensis Euonymus fimbriatus Euonymus lawsonii Euonymus benguetensis Euonymus boninensis Euonymus chenmoui Euonymus chloranthoides Euonymus fusiformis Euonymus hemsleyanus Euonymus hukuangensis Euonymus revolutus Euonymus rothschuhii Euonymus sootepensis Euonymus atropurpureus Euonymus verrucosus Euonymus americanus Euonymus latifolius Euonymus alatus Euonymus japonicus Euonymus fortunei