Inflorescences of dense sessile clusters, axillary or on reduced branches, bisexual or unisexual by abortion, always without a common involucre, but often with ciliate or bristle-bearing bracts.
Male flowers few together, sessile; perianth boat-shaped; stamen single, inflexed; rudimentary ovary absent.
Female flowers sessile; perianth and staminodes absent; ovary erect, stigma sessile, filiform.
Leaves alternate, petiolate, serrate, dentate or entire; stipules lateral, free.
Slender annual herbs, monoecious or dioecious by abortion.
Achene with a narrow, slightly unilateral wing.
Cystoliths dot-like.