Plants small to medium sized, frequently gregarious. Stems irregularly branched, occasionally tomentose; central strand absent or weakly differentiated. Leaves oblong, elliptic, narrowly spathulate or occasionally lanceolate; apex broadly acute to rounded, mucronate to hair-pointed, occasionally cucullate; margins plane to weakly recurved on one or both sides, entire; costa single, subpercurrent to long-excurrent, awn smooth, hyaline; distal laminal cells more or less quadrate, papillose on one or both exposed surfaces with several large, branching or C-shaped papillae; marginal cells sometimes longer proximally, forming a very weak border; basal cells long-rectangular, thin-walled, generally smooth, cross walls frequently distinctly colored. Sexual condition autoicous. Seta elongate, smooth, brown to red or dark red. Capsule generally long-exserted or just emergent, erect to inclined, cylindric, smooth or distinctly furrowed; gymnostomous or peristomate, stomata few; peristome single, double or absent, exostome frequently reduced to small, irregular projections or teeth long, narrow, papillose; operculum usually conic-rostrate. Calyptra smooth or papillose distally or frequently only on rostrum, entire, fringed or lacerate below, fringe small or well developed. Spores generally large, ornamentation papillose to warty, or ridged.
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