Erythroxylaceae Kunth

Coca family (en)

Family

Angiosperms > Malpighiales

Characteristics

Glabrous trees or shrubs, sometimes with milky or coloured sap. Leaves alternate or opposite (Aneulophus), simple, entire; stipules present, intrapetiolar or interpetiolar (Aneulophus), usually caducous, with colleters. Inflorescences subumbelliform cymes (Pinacopodium), fascicles, or flowers solitary, axillary. Flowers bisexual or rarely unisexual (not in Australia), actinomorphic, pedicellate or sessile (not in Australia), often distylous. Sepals rarely 4 (not in Australia) or 5, basally fused or free, persistent, valvate or imbricate in bud. Petals 5 or rarely 4 (not in Australia), free, caducous, imbricate in bud, usually with a bifid nectariferous appendage on inner face, rarely without (not in Australia). Stamens 10 in 2 series, rarely 12 (not in Australia); filaments connate at base into a cup or tube; anthers basifixed, tetrasporangiate, longitudinally dehiscent. Ovary superior, usually 3-, rarely 2-(not in Australia) locular with only 1 locule fertile, or 3 or 4 fertile locules (Aneulophus), with free or fused styles; ovules 1 per locule, rarely 2 (Aneulophus); placentation axile. Fruit a drupe or a septicidal capsule (Aneulophus). Seeds 1 or 2–3 (not in Australia), arillate (Aneulophus) or not.
More
Trees, shrubs or subshrubs, glabrous. Leaves alternate or rarely opposite, simple, entire, pinnately nerved; stipules ± united, intrapetiolar or rarely interpetiolar. Flowers axillary, solitary, in fascicles or rarely in pedunculate inflorescences, regular, hermaphrodite or unisexual, heterostylous. Sepals (4–)5, valvate, ± united. Petals (4–)5, with contorted aestivation, free, clawed, soon falling, usually with a ligulate nectariferous appendage inside. Stamens 10, all fertile, the filaments united at the base to form a shallow or urn-shaped cup, sometimes glandular; anthers basifixed, dehiscing longitudinally. Ovary superior, 2–3(–4)-locular, the locules with 1 or rarely 2 pendulous ovules; styles 2–3(–4), free or ± united, with club-shaped, capitate or rarely acute stigmas. Fruit a 1-seeded fleshy drupe or rarely a 3(–4)-locular, 3(–4)-seeded capsule dehiscing longitudinally. Seeds with no aril and no or very little endosperm; embryo straight with flat or semiconvex cotyledons
Shrubs or trees. Stipules intrapetiolar. Leaves alternate or rarely opposite, simple; leaf blade margin entire. Flowers usually bisexual, in axillary fascicles or cymes, regular, 5-merous, often heterostylous. Sepals 5, basally connate, with imbricate or valvate lobes, persistent. Petals 5, distinct, imbricate, usually with a scale on inner face at base. Stamens 5, 10, or 20, 1-or 2-verticillate; filament bases usually connate into a tube; anthers elliptic, 2-celled, with longitudinal slits. Ovary superior, connected with 3-5 carpels, 3-5-locular, with 1(or 2) axile; ovules pendulous, anatropous to hemitropous, placentation axile; styles 1-3 or 5, distinct or somewhat connate; stigmas oblique. Fruit a capsule or a 1-seeded drupe. Seeds with straight embryo and copious (rarely absent) endosperm.
Ovary superior, 2–3 (4)-locular, with each loculus 1-ovulate (or rarely 2-ovulate); ovules pendulous; styles 2–3 (4), free or ± united, with stigmas clavate to obliquely capitate or depressed-capitate rarely acute
Ovary of 3 carpels, 3-celled, mostly two of the cells sterile, fertile cells 1–2-ovuled; ovules pendulous; styles 3, free or more or less connate; stigmas oblique, depressed-capitate or clavate
Stamens 5 + 5, all fertile, with filaments united at the base to form a deep urceole or shallow rarely glandiferous cup; anthers basifixed, dehiscing longitudinally
Leaves alternate (or rarely opposite), simple, entire, penninerved, with stipules ± united and intrapetiolar (or rarely interpetiolar)
Flowers axillary, solitary or in fascicles (rarely pedunculate), actinomorphic, bisexual or rarely subdioecious, heterostylic
Seeds not compressed, exarillate, without or with scanty endosperm; embryo straight, with flat or semiconvex cotyledons
Petals (4) 5, contorted in bud, free, unguiculate, caducous, usually with a ventral ligule-like nectariferous appendage
Leaves alternate, rarely opposite, simple, entire; stipules intrapetiolar, rarely extrapetiolar, often caducous
Stamens 10, in 2 series, more or less connate at the base; anthers ellipsoid, 2-celled, opening lengthwise
Fruit a 1-seeded fleshy drupe (or rarely a 3 (4)-locular 3 (4)-seeded capsule dehiscing longitudinally)
Flowers fasciculate, hermaphrodite, rarely subdioecious, hypogynous, actinomorphic
Petals 5, free, deciduous, imbricate, mostly ligulate on the inside
Fruit drupaceous; seeds with or without endosperm; embryo straight
Calyx persistent, campanulate, lobes 5, imbricate
Trees, shrubs or shrublets, glabrous
Sepals (4) 5, valvate, ± united
Trees, shrubs, or undershrubs
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Foliage retention deciduous
Sexuality hermaphrodite
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Usage

The drug cocaine is obtained from Erythroxylum coca and E. novogranatense.
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Cultivation

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Images

Erythroxylaceae unspecified picture

Distribution

Erythroxylaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:30006363-2
WFO ID wfo-7000000221
COL ID 9VH
BDTFX ID -
INPN ID 597856
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Erythroxylaceae

Lower taxons

Aneulophus Nectaropetalum Pinacopodium Erythroxylum