Erythroxylum P.Browne

Coca (en)

Genus

Angiosperms > Malpighiales > Erythroxylaceae

Characteristics

Glabrous trees or shrubs; twigs compressed, usually becoming terete; sap sometimes yellow or reddish, bitter. Leaves simple, entire, minute to large, al-ternate-distichous, deciduous often several times a year, ptyxis involute often leaving lines or a distinctly textured areole on the leaf undersides, venation pin-nate, the midvein prominent, often excurrent beneath, the lateral veins mostly numerous, anastomosing irregularly with reticulate minor venation both within and beyond the arc of the lateral veins; petioles basally articulated; stipules mostly fused forming a single structure distal to the petiole with 2 median ribs and in some species with numerous striations, sometimes setose or fimbriate, on new growth or near inflorescences sometimes appearing without a leaf (ramenta) or with the leaf reduced to a minute cylindrical stub. Inflorescence mostly fas-ciculate, appearing as the leaves mature, subtended by bracts which resemble the stipules; the pedicels mostly angled, articulated near the base and accom-panied by paired bractlets (prophylls). Flotwers mostly perfect, sometimes subdioecious, heterostylous with the styles and filaments alternating in length; calyx 5-lobed, sometimes wing-angled, the sepals imbricate or valvate, some-times to a sharp point, persistent; petals free, deciduous, mostly white, the bot-tom third demarcated from the upper portion, mostly with 2 more or less fused petaloid appendages arising from the horizontal demarcation, often with a nec-tary situated between the appendages and petal lamina and median to the line of demarcation; stamens 10, in 2 series, the outermost alternate with the petals, the filaments basally fused into a tube, the innermost sometimes ventral to the truncate or dentate rim of the tube, the outermost arising directly from the tube, the anthers 2-loculed, rotund, basifixed, longitudinally dehiscent; ovary ellip-soid or ovoid, mostly slightly truncate, 3-loculed, only one locule ovule-bearing, this with one (?rarely 2) pendulous, epitropous, anatropous ovule. Fruit a small, red, baccate drupe, the endocarp hard; seed shiny, endosperm present or want-ing, sometimes copious.
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Shrubs or trees. Youngest branchlets compressed, older branches terete; base of the lateral twigs often provided with small distichous 'bracts' (ramenta) sometimes also occurring between the leaves. Leaves simple, alternate (distichous), entire, involute in bud, the margins leaving a more or less permanent trace as 2 longitudinal lines on the upper leaf surface ('areolate'). Stipules mostly entirely connate, rarely bifid, intrapetiolar, often bicarinate, sometimes emarginate or 2-toothed at the apex, long-persistent or early caducous, inserted ± semi-amplexi-caulous and leaving a distinct, mostly oblique scar. Flowers axillary, solitary or in clusters, often dimorphous, or even 3-4-morphous, 5-merous, actinomorphic, bisexual. Pedicels more or less thickened, often only under the calyx, provided with 2 bracteoles at the base. Calyx persistent, campanulate, (in Mal.) ± halfway divided into 5 lobes imbricate in bud. Petals 5, free, caducous, alternating with the calyx lobes, quincuncial in bud, nearly always provided with an emarginate or 3-lobed ligule inserted on the apex of the claw of the petal. Stamens 10, in two whorls of 5, persistent; filaments towards the base connate into a staminal tube often with a toothed margin; anthers ellipsoid, basifixed, cordate at the base, 2-celled, opening lengthwise, latrorse. Ovary (1-)3-celled, each cell with 1 ovule, normally only 1 cell fertile, but the other empty cells sometimes distinctly enlarged in fruit; styles 3, erect, free or partly connate or stigmas ± sessile; stigmas flattened (often oblique) or (in extra-Mal. spp.) clavate, blunt or rarely acute. Ovules pendulous, anatropous, with a ventral raphe. Fruit a drupe. Seed with or without endosperm; embryo oblong, erect; cotyledons flat to plano-convex; no plumule, but a distinct radicle.
Trees, shrubs or shrublets, the young shoots often ± markedly compressed. Leaves petiolate, alternate, involute in bud; stipules intrapetiolar, ± united, persistent or partially to completely deciduous. Flowers axillary, in fascicles or solitary, heterostylous. Sepals 5, valvate, triangular, connate and broadened at the base, ± coriaceous. Petals white to yellowish cream, free, clawed; nectary-appendage large, usually exceeding the calyx, entire or 2–3-lobed. Stamens with the filaments equal or unequal and then those opposite the sepals shorter, united at the base to form a deep ± persistent cup with entire or ± toothed margin. Ovary (2–)3-locular, the locules with solitary ovules; styles free or partially united, spreading; stigmas obliquely capitate or rarely acute. Fruit a 1-seeded drupe with the sterile locule ± apparent. Seed oblong-ellipsoid; endosperm usually fleshy.
Shrubs or small trees, usually glabrous. Stipules intrapetiolar, often imbricating on short branches. Leaves alternate, often subdistichous, simple. Flowers axillary, solitary or fascicled, small, often heterostylous. Sepals 5(or 6), usually basally connate. Petals 5, white or pink, clawed, usually with a ligule on inner face at base, imbricate. Stamens 10; filament usually basally connate into a glandular or nonglandular tube. Ovary 3(or 4)-locular, with 1(or 2) ovules in single fertile locule; styles distinct or connate; stigmas oblique. Drupe 1-seeded. Seed with a thin testa; endosperm variable in quantity or absent; embryo straight.
Leaves petiolate, alternate, simple, entire, involute when young, with venation varying in prominence and degree of reticulation; stipules intrapetiolar, ± united, persistent or partially to completely deciduous or caducous.
Stamens 5 + 5, with filaments equal or unequal (antisepalous shorter than antipetalous), united in the lower part to form a deep cup with margin entire or ± denticulate.
Petals white to cream-yellow, free, unguiculate; nectary appendage (ligule) large, usually exceeding the calyx, entire or 2–3-lobed.
Ovary (2) 3-locular with 1-ovulate loculi; styles free or partially united, divergent; stigmas obliquely capitate, rarely acute.
Sepals 5, valvate, triangular, connate and broadened at the base, ± coriaceous.
Trees, shrubs or shrublets, with young shoots usually ± compressed.
Petals as many as sepals, imbricate, with a scale on the inner side
Flowers axillary, in fascicles or solitary, heterostylic.
Fruit a 1-seeded drupe, with sterile loculi ± apparent.
Seed oblong-elliptic, usually with fleshy endosperm.
Flowers small, pedicellate, in axillary fascicles
Leaves alternate, entire
Stipules, intrapetiolar
Shrubs or small trees
Ovary 3-4 celled
Sepals usually 5
Styles 3 or 4
Fruit a drupe
Stamens 10
Pending.
Life form perennial
Growth form tree
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
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Environment

The Malaysian spp. are confined to the substage of the primary rain-forest up to c. 1600 m, obviously avoiding areas subject to a dry season; of E. cuneatum only two specimens have been collected in the seasonal area of the Lesser Sunda Islands, viz on an islet of the Kangean group and on Sumbawa. In Africa and Australia some xerophytic species are known under arid climatic conditions.E. cuneatum is often found in the coastal forest along the beach or behind the mangrove as a small tree.
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Hardiness (USDA) 8-12

Usage

Uses. E. coca Lamk and E. novogranatense (Morris) Hieron. are very important because of the high content in valuable alkaloids found in their leaves. Cocaine is used nowadays medicinally throughout the world as a stimulant and as an anaesthetic, specially in eye-surgery.In Malaysia only E. novogranatense is grown for coca. In trade it is generally known as 'Java coca' or 'Truxillo coca'.E. cuneatum (Miq.) Kurz furnishes a hard and durable, pink-coloured timber, which is through its often small dimension and quantity of minor importance.
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The South American species Erythroxylum coca, is the source of the drug cocaine used in medicinal products and illegal trade. 
Uses medicinal timber
Edible -
Therapeutic use -
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Cultivation

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Images

Erythroxylum unspecified picture

Distribution

Erythroxylum world distribution map, present in Australia, China, Fiji, Micronesia (Federated States of), Indonesia, Madagascar, Malaysia, Panama, Solomon Islands, United States of America, Samoa, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:24933-1
WFO ID wfo-4000014016
COL ID 62S56
BDTFX ID -
INPN ID 627958
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Sethia Erythroxylum

Lower taxons

Erythroxylum acrobeles Erythroxylum amazonicum Erythroxylum amplifolium Erythroxylum amplum Erythroxylum ampullaceum Erythroxylum anceps Erythroxylum andrei Erythroxylum anguifugum Erythroxylum apiculatum Erythroxylum areolatum Erythroxylum argentinum Erythroxylum arrojadoi Erythroxylum badium Erythroxylum bezerrae Erythroxylum bicolor Erythroxylum bradeanum Erythroxylum brevipes Erythroxylum buxifolium Erythroxylum buxus Erythroxylum caatingae Erythroxylum cambodianum Erythroxylum campestre Erythroxylum campinense Erythroxylum capitatum Erythroxylum cincinnatum Erythroxylum citrifolium Erythroxylum clarense Erythroxylum coelophlebium Erythroxylum columbinum Erythroxylum cuneatum Erythroxylum cuneifolium Erythroxylum densum Erythroxylum discolor Erythroxylum ecarinatum Erythroxylum emarginatum Erythroxylum ferrugineum Erythroxylum firmum Erythroxylum fischeri Erythroxylum flavicans Erythroxylum foetidum Erythroxylum gerrardii Erythroxylum gracilipes Erythroxylum grandifolium Erythroxylum grisebachii Erythroxylum guanchezii Erythroxylum guatemalense Erythroxylum hamigerum Erythroxylum haughtii Erythroxylum hildebrandtii Erythroxylum hondense Erythroxylum horridum Erythroxylum hypericifolium Erythroxylum hypoleucum Erythroxylum impressum Erythroxylum incrassatum Erythroxylum jamaicense Erythroxylum lanceum Erythroxylum leptoneurum Erythroxylum ligustrinum Erythroxylum longipes Erythroxylum loretense Erythroxylum lygoides Erythroxylum macrocalyx Erythroxylum macrochaetum Erythroxylum macrophyllum Erythroxylum magnoliifolium Erythroxylum mangorense Erythroxylum mannii Erythroxylum maracasense Erythroxylum martii Erythroxylum mattos-silvae Erythroxylum membranaceum Erythroxylum mexicanum Erythroxylum microphyllum Erythroxylum mikanii Erythroxylum minutifolium Erythroxylum monogynum Erythroxylum nitidulum Erythroxylum nobile Erythroxylum nossibeense Erythroxylum novocaledonicum Erythroxylum novogranatense Erythroxylum obtusifolium Erythroxylum occultum Erythroxylum ochranthum Erythroxylum opacum Erythroxylum oreophilum Erythroxylum orinocense Erythroxylum paraguariense Erythroxylum parvistipulatum Erythroxylum passerinum Erythroxylum patentissimum Erythroxylum pauferrense Erythroxylum pedicellare Erythroxylum pelleterianum Erythroxylum pervillei Erythroxylum petrae-caballi Erythroxylum platycladum Erythroxylum plowmanianum Erythroxylum plowmanii Erythroxylum polygonoides Erythroxylum popayanense Erythroxylum pruinosum Erythroxylum pulchrum Erythroxylum pungens Erythroxylum raimondii Erythroxylum reticulatum Erythroxylum rignyanum Erythroxylum rotundifolium Erythroxylum rufum Erythroxylum ruizii Erythroxylum santosii Erythroxylum sarawakanum Erythroxylum schomburgkii Erythroxylum sechellarum Erythroxylum steyermarkii Erythroxylum strobilaceum Erythroxylum suberosum Erythroxylum subglaucescens Erythroxylum subracemosum Erythroxylum subrotundum Erythroxylum subsessile Erythroxylum substriatum Erythroxylum urbanii Erythroxylum vernicosum Erythroxylum virgultosum Erythroxylum williamsii Erythroxylum zambesiacum Erythroxylum zeylanicum Erythroxylum jaimei Erythroxylum longisetulosum Erythroxylum timothei Erythroxylum dillonii Erythroxylum gentryi Erythroxylum splendidum Erythroxylum acuminatum Erythroxylum affine Erythroxylum alaternifolium Erythroxylum barbatum Erythroxylum bequaertii Erythroxylum betulaceum Erythroxylum carthagenense Erythroxylum cassinoides Erythroxylum cataractarum Erythroxylum catharinense Erythroxylum compressum Erythroxylum confusum Erythroxylum cordato-ovatum Erythroxylum coriaceum Erythroxylum corymbosum Erythroxylum couveleense Erythroxylum cuspidifolium Erythroxylum daphnites Erythroxylum deciduum Erythroxylum distortum Erythroxylum divaricatum Erythroxylum domingense Erythroxylum dumosum Erythroxylum ellipticum Erythroxylum engleri Erythroxylum frangulifolium Erythroxylum gaudichaudii Erythroxylum glaucum Erythroxylum glaziovii Erythroxylum kapplerianum Erythroxylum kochummenii Erythroxylum laetevirens Erythroxylum lanceolatum Erythroxylum lancifolium Erythroxylum laurel Erythroxylum leal-costae Erythroxylum lenticellosum Erythroxylum lindemanii Erythroxylum lineolatum Erythroxylum moonii Erythroxylum mucronatum Erythroxylum myrsinites Erythroxylum myrtoides Erythroxylum nelson-rosae Erythroxylum ovalifolium Erythroxylum oxycarpum Erythroxylum oxypetalum Erythroxylum pachyneurum Erythroxylum pacificum Erythroxylum panamense Erythroxylum revolutum Erythroxylum rimosum Erythroxylum roigii Erythroxylum roraimae Erythroxylum rosuliferum Erythroxylum simonis Erythroxylum sinense Erythroxylum sphaeranthum Erythroxylum spruceanum Erythroxylum squamatum Erythroxylum stipulosum Erythroxylum subumbellatum Erythroxylum tenue Erythroxylum tianguanum Erythroxylum tortuosum Erythroxylum tucuruiense Erythroxylum ulei Erythroxylum vaccinifolium Erythroxylum vasquezii Erythroxylum glazioui Erythroxylum pictum Erythroxylum platyclados Erythroxylum delagoense Erythroxylum cuneifolioides Erythroxylum umbrosum Erythroxylum nordestinum Erythroxylum rhodappendiculatum Erythroxylum sobraleanum Erythroxylum stenopetalum Erythroxylum tapacuranum Erythroxylum ayrtonianum Erythroxylum cogolloi Erythroxylum riparium Erythroxylum umbu Erythroxylum angelicae Erythroxylum riverae Erythroxylum acutum Erythroxylum banaoense Erythroxylum davidii d'arcy Erythroxylum gonoclados Erythroxylum gracile Erythroxylum leandrianum Erythroxylum lofgrenii Erythroxylum macrocarpum Erythroxylum mamacoca Erythroxylum mogotense Erythroxylum x peyritschii Erythroxylum seyrigi Erythroxylum striiflorum Erythroxylum striolatum Erythroxylum undulatum Erythroxylum vaginatum Erythroxylum mocquerysii Erythroxylum armatum Erythroxylum australe Erythroxylum bangii Erythroxylum baracoense Erythroxylum boinense Erythroxylum boivinianum Erythroxylum brennae d'arcy Erythroxylum ruryi Erythroxylum schunkei Erythroxylum shatona Erythroxylum socotranum Erythroxylum sparsiflorum Erythroxylum xerophilum Erythroxylum dekindtii Erythroxylum elegans Erythroxylum nummularia Erythroxylum pauciflorum Erythroxylum pyrifolium Erythroxylum sideroxyloides Erythroxylum annamense Erythroxylum fimbriatum Erythroxylum coca Erythroxylum havanense Erythroxylum laurifolium