Erythroxylum novogranatense (D.Morris) Hieron.

Colombian coca (en)

Species

Angiosperms > Malpighiales > Erythroxylaceae > Erythroxylum

Characteristics

Bush or shrub c. 1-3 m. Lenticels on the branches minute. Leaves abundant along the twigs, not soon falling, obovate-oblong, bright green above, paler and glaucous beneath, c. (2-)3-6(-7) by 1-3 cm; rounded or sometimes emarginate, always with a mucronate tip or notch, attenuate at the base; midrib rather prominent beneath, specially at the base, nerves numerous, on both surfaces equally distinct, venation delicately anastomosing, central part included between the areolation lines slightly concave and of a paler colour on both surfaces; petiole thin, c. 3-7 mm. Stipules green, triangular, shorter or half as long as the petiole, c. 2.5-4 by 2-3 mm, slightly bicarinate, persistent, on old branches brown, top emarginate, 2-toothed, margin more or less fimbriate. Ramenta none or occasionally a few. Flowers in clusters of 1-4(-8). Bracteoles green, deltoid, acuminate, scarious, up to c. 2.5 by 1 mm, margin delicately fimbriate. Pedicels thickened, c. 5-10 mm. Calyx green, the tube 1-1.5 mm, lobes triangular-ovate, acuminate, c. 1.5-2 by 1 mm. Petals white or greenish-white, oblong, convex c. (3.5-)4(-5) by 2 mm; ligule 3-lobed, with crisped lobes, half as long as the blade of the petal; claw c. 1/5-1/4 the length of the petal. Brachystylous flowers. Staminal tube pale green c. 1 mm. Stamens equal, filaments c. 3½-5½ mm, anthers c. 0.5 by ⅔ mm. Ovary distinctly longer than the staminal tube, c. 2 by 1 mm; styles free, c. 1 mm. Dolichostylous flowers. Staminal tube 1 mm. Stamens unequal, episepalous filaments ½-¾-1 mm, epipetalous ones 0.5-1.5-2 mm, anthers 0.75-0.5 by 0.5-0.25 and 0.5 by 0.25 mm respectively. Ovary longer than the staminal tube, c. 1½ by 1 mm; styles free, c. 2-3 mm. Drupe red, elliptic-oblong, in the fresh state 9-11 by 4-5 mm; stone narrower, pointed, obtusely trigonous, with 4-6 ribs and furrows, c. 8-10 by 3-4 mm; sterile cells hardly distinct, fertile cell almost triangular in section. Seed with 3 distinct ribs; endosperm abundant; embryo central, small, c. 6-7 by 1 mm, flattened; cotyledons lanceolate, c. 3-4 mm; radicle c. 2-3 by 0.5 mm.
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Shrubs or small trees. Stipules triangular, 1.5-3 mm, margin often entire. Leaves alternate, simple; petiole 4-7 mm; leaf blade obovate to narrowly elliptic, 1.2-4.7 × 1-1.8 cm, midvein adaxially impressed, secondary veins anastomosing into an obvious intramarginal vein in each side of midvein, base attenuate, margin entire, apex obtuse to retuse and with a mucro. Flowers solitary or fascicled, small. Pedicel to 4 mm at anthesis. Sepals 5, ca. 1.5 mm, basally connate. Petals 5, whitish yellow, ovate-oblong, 3-3.5 mm, with 2 ligules on inner face at base. Stamens 10; filament basally connate into a tube, 2-4 mm. Ovary globose to oblong, 1-3.5 mm, 3-locular, with 1 ovule in only fertile locule; styles distinct, 1-3 mm, persistent. Mature drupe red, oblong, 7-8 × ca. 3 mm, with 5 longitudinal ribs, 1-seeded. Fl. most of year but mostly Feb-Mar, fr. May-Dec.
A shrub or small tree. The leaves are simple and alternate. They are narrowly oval and 1-5 cm long by 1-2 cm wide. Flowers can occur singly or in groups. They are small and whitish yellow. The fruit are red and oblong. They are 7-8 mmlong by 3 mm wide.
Life form perennial
Growth form shrub
Growth support free-standing
Foliage retention evergreen
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 2.5 - 3.0
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway -

Environment

A tropical plant. It grows up to 1600 m altitude in the Andes. It suits the hot moist tropics. In XTBG.
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In Malaysia often grown as a hedge plant, up to 750 m. Fl.-fr. Jan.-Dec.
Not known
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses. Cultivated in Java since 1875 in the Botanic Garden at Bogor, when the first seeds were received. In 1885 after many difficulties and contradictory reports and advices an experimental plantation was started in the Tjikeumeuh Agricultural Garden at Bogor from seeds imported by the firm of Herman Linden & Co, which proved successful. Since 1886 many seeds were distributed to new plantations. The culture never gained great importance in Java; the price of the leaves rapidly decreased in the early twenties. Several estates used the species as hedges and as accessory crop. According to Corner ( Corner Ways. Trees 1 1940 220 ) cultivation was prohibited in the Malay Peninsula.From the dried leaves medicinal tinctures were prepared and by refining these alkaloids are obtained of which the main one is cocaine, which is used as a stimulant and as an anaesthetic mainly for eye-surgery. In Java a factory was built to prepare the crude alkaloid for export in order to avoid the costs of transport of the leaves to Europe ( Burkill Dict. 1 1935 952 ).
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The leaves are chewed as a stimulant. They are used in drinks.
Uses environmental use material medicinal poison social use
Edible leaves
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Plants can be grown from seeds and from cuttings.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Erythroxylum novogranatense unspecified picture

Distribution

Erythroxylum novogranatense world distribution map, present in China, Colombia, Costa Rica, Ecuador, Guadeloupe, Grenada, Honduras, Indonesia, Iceland, Martinique, Malaysia, Nicaragua, Panama, Peru, Philippines, Trinidad and Tobago, Taiwan, Province of China, Venezuela (Bolivarian Republic of), Viet Nam, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:97004-2
WFO ID wfo-0000680010
COL ID 3BFYS
BDTFX ID -
INPN ID 629673
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Erythroxylum novogranatense Erythroxylum coca var. novogranatense Erythroxylum coca var. spruceanum Erythroxylum novogranatense var. microphyllum Erythroxylum novogranatense var. novogranatense

Lower taxons

Erythroxylum novogranatense var. truxillense Erythroxylum novogranatense var. tobagense