Rhizome terete, 2-5 mm thick, brown; phyllopods 1-7 cm distant. Anatomy: vascular strands 7-10, with dark bundle sheaths, sclerenchyma strands many (60-90). Rhizome scales inserted evenly, persistent, densely set, spreading, pseudopeltate, monomorphic, deltoid, 2-5 by 1.3-2.6 mm, index 1.5-2, basal auricles rounded, margin at base sparsely set with glands, dentate but entire near the base, apex acuminate, cell walls not thickened at base of auricles, thinly clathrate in upper part, cells narrow, c. 20 in a row between the two margins at the base of the acumen, hyaline, a tuft of hairs usually present near the point of attachment. Fronds (pendent) pinnate, stipe 2-5 mm thick, 0.4-0.6 as long as the lamina, glabrous; lamina 40-70(-100) by 20-25 cm, index 2-2.8, widest at or below the middle; pinnae articulate, sessile, 11-19 by 0.5-1.5 cm, index 7.3-22, widest in the lower half, base cuneate to truncate, rarely cordate, margin serrate, apex very narrowly acute to acuminate; basal pair 2-3 cm distant from next pair, deflexed or not, apical segment conform, half to as long as longest pinnae, often lobed at the base. Venation: areoles uniserial, marginal veinlets very short or well-developed, rarely anastomosing into areoles which then lack included veinlets. Indument: 2-or 3-celled glandular hairs present, acicular hairs in varying density on veins and lamina, 2-6 cells long, scales persistent. Sori uniserial, marginal, slightly to deeply sunken, forming distinct papillae on the adaxial surface, 1.2-2.2 mm diam., receptacular scaly paraphyses deciduous, clathrate, 100-200 by 100-140 µm, 2 or more cells wide. Sporangia glabrous. Spores yellow, cristate, with short polar excrescences.
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