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Rhizome long-creeping, often glaucous; scales attenuate, clathrate, ±deciduous. Fronds monomorphic. Stipes long, distant, articulated to short phyllopodia. Lamina pinnate or (not in Australia) pinnatifid; pinnae articulated to rachis in pinnate species. Veins forked, forming a series of areoles on each side of costa; areoles with a free included veinlet branching from near the base of a main lateral vein on apical side and directed towards margin; outer veins often free or sometimes forming 1 or more similar or smaller series of areoles, with or without free included veinlets. Sori (in Australian spp.) in 1 row on each side of costa, terminal on the included veinlets of the inner series of areoles, usually ±impressed into lamina. Spores bilateral, hyaline, yellow or brownish, smooth; perispore inconspicuous or cristately folded.
Epiphytic or epilithic, occasionally terrestrial. Rhizome long-creeping, phyllopods 0.5-5 cm distant. Anatomy: ground tissue parenchymatous, sclerenchyma strands present. Rhizome scales basifixed to pseudopeltate, entirely or mostly clathrate, appressed to erect, margin dentate, apex acuminate to filiform, often with superficial hairs. Fronds monomorphic, stipitate, pinnate to pinnatifid, margin crenate to serrate. Venation with 1-several rows of areoles each with 1 included, excurrent, free veinlet; marginal row of excurrent veinlets present. Hydathodes present. Sori round, in 1-3 rows between pinna-midrib and margin, superficial to deeply sunken. Scaly paraphyses present, long-stalked, with basifixed or peltate, clathrate blade.
Some veins anastomosing
