Rhizome terete, 5-11 mm thick, brown; phyllopods 1.5-2.5 cm distant. Anatomy: vascular strands 14-19, with dark bundle sheaths, sclerenchyma strands many (70-100). Rhizome scales inserted evenly, deciduous, densely set, appressed, pseudopeltate, monomorphic, deltoid, 2.5-4 by 0.7-1.3 mm, index 2.8-3.6, basal auricles rounded, margin at the base sparsely set with glands, dentate, teeth all equally long or slightly shorter at base, apex acuminate to filiform, cell walls equally clathrate throughout, superficial hairs present, spread over the entire acumen. Fronds pinnate, stipe 3.5-5 mm thick, 0.2-0.6 as long as the lamina, glabrous; lamina 38-99(-200) by 18-38(-45) cm, index 2.1-4.4, widest at the base; pinnae articulate, shortly petiolate, 12.5-20.5 by 2-3 cm, index 6.2-6.8, base shortly angustate, margin entire to crenate, apex acuminate; basal pair 2-6 cm distant from next pair and equally long, not deflexed, apical segment conform, slightly longer than longest pinnae/segment. Venation: areoles 3-or 4-serial, marginal veinlets up to 1/3 as long as the areole, sim-ple or forked, frequently anastomosing. Indument: 2-celled glandular hairs present, acicular hairs absent to dense all over the lamina, 2-5 cells long, scales deciduous. Sori uniserial, costal, deeply sunken, 0.8-1 mm diam., receptacular scaly paraphyses persistent, clathrate, 230-250 by 180-200 µm, 3 or more cells wide. Sporangia glabrous. Spores yellow, cristate, with short polar excrescences.
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Rhizome not chalky white. Scales caducous, dark brown, 1.2-4 mm long, 0.7-1.5 mm wide; base broad and rounded; margins with fine rigid teeth; apex narrow, hair-like. Fronds to 1-2 m or more long. Stipe 8-34 cm long, scaly at base; scales as for rhizome but usually longer and narrower. Lamina 1-pinnate, becoming pendulous. Pinnae narrowly ovate to oblong, commonly 3-5 cm apart, 3-22.5 cm long, 0.7-3 cm wide, thin, firm, clothed when young with soft pale hairs, glabrescent apart from some small scattered scales on rachis and lower surface of costae; base equally or unequally cuneate; margins ±entire to shallowly dentate; apex acuminate, acute. Veins usually forming 3-4 series of areoles between costa and margin; areoles mostly with free included veinlets. Sori solitary in each of the innermost areoles, surrounded by a ring of dark brown dentate scale-like paraphyses; sporangia immersed in distinct depressions. Spores 30-47.5 µm long, 15-30 µm wide.
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