Rhizome terete, 3.5-4.5 mm thick, brown; phyllopods 1.5-10 cm distant or more. Anatomy: vascular strands 11-14, with dark bundle sheaths, sclerenchyma strands many, up to 50-60. Rhizome scales inserted evenly or on protrusions, deciduous, densely set, rather suddenly contracted from a wide base to a squarrose, slightly channeled acumen, basifixed or pseudopeltate, monomorphic, deltoid, 2.1-7.3 by 0.4-2.1 mm, index 3.5-2.2, basal auricles rounded, margin of the acumen sparsely set with glands, base entire to dentate, acumen entire or denticulate, apex filiform, cell walls clathrate throughout, more strongly thickened in the central part of the acumen, sometimes forming a light pseudomidrib, superficial hairs absent. Fronds pinnatifid, stipe 3-4 mm thick, 0.6-0.8 as long as the lamina, hairy; lamina 20-45 by 18-28 cm, index 1.1-1.6, widest near the base; segments 10-15 by 1.3-1.5 cm, index 7.7-10, base of lowest pinnae cordate on basiscopic side, margin entire to crenate, apex acuminate, basal pair 1.5-4.5 cm distant from next pair and equally long, basal 1 or 2 pairs rather strongly deflexed, apical segment not sharply distinct. Venation: areoles 1-or 2-serial, marginal veinlets up to half as long as the areole, simple or forked, frequently anastomosing to form areoles without included veinlets. Indument: 2-4-celled glandular hairs present, acicular hairs present on upper surfaces of midrib and costae, sometimes also on lower surfaces and on the lamina, 2-5 cells long, scales deciduous or sometimes persistent. Sori uniserial, costal, superficial, 1.5-2 mm diam., receptacular scaly paraphyses persistent, clathrate, 140-180 by 90-140 µm, 2 cells wide. Sporangia glabrous. Spores yellow, reticulate, without polar excrescences.