Rhizome terete, 3.5-5 mm thick, slightly glaucous to white-waxy; phyllopods 1.5-6.5 cm distant. Anatomy: vascular strands 8-15, without dark bundle sheaths, sclerenchyma strands 40-90. Rhizome scales inserted evenly, blackish, densely set, dimorphic; large scales often caducous, erect, pseudopeltate, deltoid, 1.7-3.9 by 0.5-1.1 mm, index 3.4-3.6, basal auricles rounded, irregularly dentate, acumen contracted, apex filiform; margin densely set with glands, dentate, teeth longer at base, cell walls clathrate throughout, walls thicker in the acumen towards apex, cells hyaline, superficial hairs absent; small scales brownish, abundant, persistent, irregularly shaped, 0.06-0.1 mm diam. Fronds pinnate, stipe 2-3.5 mm thick, 0.2-0.7 as long as the lamina, glabrous; lamina 20-60(-70) by 8-15(-20) cm, index 2.5-4, widest at the middle or equally wide throughout; pinnae articulate, sessile at the base of the lamina, to adnate and decurrent on the rachis in at least the upper 1/3 of the lamina, 8-11 by 1-1.3 cm, index 8-8.5, base cordate or auricled, rarely truncate or cuneate, margin entire to crenate, apex narrowly acute to acuminate; basal pair 1.5-4.5 cm distant from next pair and equally long, uppermost pinnae not strongly reduced, apical segment conform, sometimes lobed at the base, as long as or slightly longer than longest pinnae. Venation: areoles uniserial, marginal veinlets to half as long as the areole, simple or forked, frequently anastomosing to form empty areoles. Indument: 2-5-celled glandular hairs present, acicular hairs absent, scales deciduous or sometimes persistent. Sori uniserial, medial, superficial, 0.8-1.5 mm diam., receptacular scaly paraphyses persistent, clathrate, peltate, stellate or deltoid, 150-250 by 140-250 µm, 3 or more cells wide. Sporangia glabrous. Spores yellow, cristate, without polar excrescences.
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