Rhizome terete, 5-6 mm thick, brown; phyllopods 2-5 cm distant. Anatomy: vascular strands 11-13, with dark bundle sheaths, sclerenchyma strands very many (> 100). Rhizome scales inserted evenly, deciduous, densely set, spreading, pseudopeltate, monomorphic, deltoid, 1.4-4.4 by 0.5-0.9 mm, index to c. 9, basal auricles rounded, margin without glands, dentate, teeth longer at base, apex acuminate to filiform, cell walls equally strongly clathrate throughout, cells yellowish, superficial hairs sometimes present. Fronds pinnate; stipe 2-4.5 mm thick, 0.3-0.5 a as long as the lamina, glabrous; lamina 26-69 by 11-23 cm, index 2.2-3.1, widest at or near the base; pinnae articulate, sessile, 6-13 by 1-1.8 cm, index 6-7.3, base truncate, margin serrate, apex obtuse to narrowly acute or acuminate; basal pair 3-4 cm distant from next pair and slightly shorter or equally long, not deflexed, apical segment conform, sometimes with two small separate auricles at base, half to almost as long as longest pinnae. Venation: areoles uni-to biserial, second row with or without included free veinlets, marginal veinlets up to 1/3 as long as the areole, simple or forked, occasionally anastomosing. Indument: 2-celled glandular hairs present, acicular hairs absent, scales deciduous or sometimes persistent. Sori uniserial, costal, deeply sunken, forming very distinct narrow, often caducous, papillae on upper surface, 0.4-0.5 mm diam., receptacular scaly paraphyses deciduous, clathrate, 100 by 65 µm, 2 cells wide. Sporangia glabrous. Spores brownish, cristate, with short polar excrescences.