Rhizome terete, 5-7.5(-10) mm thick, white-waxy; phyllopods 3-5 cm distant. Anatomy: vascular strands 13-15, with dark bundle sheaths, sclerenchyma strands many (30-50). Rhizome scales inserted on short protrusions, deciduous, densely set, spreading or erect, basifixed to pseudopeltate, monomorphic, deltoid, 4-5 by 1-1.3 mm, index 3.8-4.1, basal auricles rounded, entire to dentate, acumen entire to dentate, apex narrow but not filiform, cell walls clathrate throughout, cells about isodiametric to elongated, hyaline, margin densely set with 1-or 2-celled glands, superficial glandular indument sporadically present. Fronds pinnate, stipe 2.5-5 mm thick, 0.4-0.6 as long as the lamina, glabrous; lamina 26-150 by 8-31 cm, index 0.4-0.6, widest near the base or equally wide throughout; pinnae articulate, sessile, 10-14 by 0.8-1.2 cm, index 11.7-12.5, adnate in upper part of the frond, base truncate, cordate or auricled, margin crenate, apex narrowly acute; basal pair 1.5-3.6 cm distant from next pair, sometimes shorter, sometimes deflexed, apical segment not sharply distinct. Venation: areoles uniserial, marginal veinlets to nearly as long as the areole, simple or forked, sometimes anastomosing. Indument: 2-celled glandular hairs present, acicular hairs present in variable densities all over the lamina, 2-8 cells long, scales persistent. Sori uniserial, medial, slightly sunken, 1.5-2 mm diam., receptacular scaly paraphyses persistent, peltate, stellate, clathrate, 200-240 by 180-200 µm. Sporangia occasionally (15% of all sporangia) with a 2-or 3-celled simple glandular hair. Spores yellow, cristate, without polar excrescences.
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