Helicia Lour.

Genus

Angiosperms > Proteales > Proteaceae

Characteristics

Trees, rarely shrubs. Leaves mostly spiral, rarely subopposite or 3-4-verticillate, simple, entire or dentate (serrate), sessile or petiolate. Racemes simple, many-flowered, axillary or ramiflorous, very rarely subterminal. Flowers bisexual. Bracts small, ovate-acuminate to subulate, in Mal. spp. subpersistent or caducous. Bracteoles minute. Pedicels mostly in twos, or solitary, free or connate in their lower part, rarely entirely connate. Perianth-tubs (in Mal. spp.) straight, slender, with a clavate to subglobose limb, somewhat swollen at the base, limb straight, segments 4, becoming free and revolute in anthesis. Stamens 4, nearly sessile, inserted at the inner base of the limb; anthers oblong, connective apiculate; exine smooth. Ovary ovoid or globose, sessile; ovules 2, anatropous, ascendent, inserted at the base of the ovary or laterally attached to the wall on the ventral suture. Style slender, clavate towards the apex; stigma punctiform. Disk glands free (spaced or approximate) or ± connate into a cup. Fruit nut-or drupelike; pericarp mostly thick-coriaceous and rather hard when dry, when fresh apparently its outer part sometimes somewhat fleshy, the inner part drier and hard, rarely with a clear differentiation in endocarp and exocarp, mostly indehiscent, sometimes very tardily splitting irregularly along the ventral (raised or sulcate) suture when dry. Seeds 1-2, subglobose respectively hemispheric. Testa thin. Cotyledons fleshy, rugulose in the upper part.
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Trees, rarely shrubs. Leaves alternate, rarely ± opposite or ± whorled, petiolate or subsessile, simple, margin entire or serrate. Inflorescence axillary, ramiflorous, rarely subterminal, a raceme of flower pairs, many-flowered. Flowers bisexual; bracts of flowers and flower pairs small, ovate-acuminate to subulate, rarely foliaceous, caducous, rarely persistent. Flower pairs sessile (“pedicels free”) or with a basal common peduncle (“pedicels connate”). Perianth tube straight, slender, limb clavate to ± globose, segments free and revolute at anthesis. Stamens subsessile, attached slightly below limb; anthers oblong; connective apiculate. Pollen grains triporate, exine scabrate-granulate. Hypogynous glands 4, free or connate into a ring or cup. Ovary sessile; ovules 2, anatropous, inserted at base of locule or laterally on wall. Style slender; stigma terminal, punctiform. Fruit usually indehiscent, although sometimes splitting along ventral suture; pericarp mostly thick and leathery. Seeds 1 or 2, ± globose to hemispheric; testa thin; cotyledons fleshy, apically rugulose.
Trees or shrubs. Branchlets terete. Leaves simple, spiral (rarely opposite); margin entire or variously toothed. Conflorescence raceme-like, terminal or subterminal, axillary, ramiflorous or rarely cauliflorous. Involucral and floral bracts small to minute, rarely persisting. Flowers actinomorphic, bisexual or andromonoecious. Receptacle straight. Perianth straight; tube slender; limb slightly expanded; tepals separate and revolute at anthesis. Hypogynous glands 4, free or fused into a cup. Stamens 4, adnate at limb base; filaments short. Ovary sessile, glabrous or tomentose; style straight; pollen presenter fusiform. Fruit drupe-like, indehiscent or splitting irregularly. Seed 1, rarely 2, globose or ovoid.
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Environment

Most species are confined to the substage of the non-seasonal rain-forest, though some are apparently adapted to a more or less seasonal climate. None is known to occur gregariously. Though most species occur in the montane zone between 500 and 2400 m, the total range is from sea-level to c. 3350 m on Mt Kinabalu, 2600 m in Central Celebes, and 2650 m in Java.
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Hardiness (USDA) 8-12

Usage

Uses. Timber is mostly of small size and specimens occur scattered so that the commercial value of this genus is of minor importance.
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Cultivation

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Images

Helicia unspecified picture

Distribution

Helicia world distribution map, present in Argentina, Australia, China, Cameroon, Indonesia, India, Japan, Malaysia, and Papua New Guinea

Identifiers

LSID urn:lsid:ipni.org:names:32922-1
WFO ID wfo-4000017216
COL ID 62TL3
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Cyanocarpus Helicia

Lower taxons

Helicia attenuata Helicia celata Helicia celebica Helicia ceylanica Helicia clemensiae Helicia coeruleopurpurea Helicia commutata Helicia ferruginea Helicia finisterrae Helicia forbesiana Helicia fuscotomentosa Helicia glabriflora Helicia grandifolia Helicia paucinervia Helicia peekelii Helicia peltata Helicia petelotii Helicia robusta Helicia rostrata Helicia rufescens Helicia saruwagedica Helicia saurauioides Helicia schlechteri Helicia serrata Helicia teysmanniana Helicia torricellensis Helicia uganensis Helicia vanroyenii Helicia versteeghii Helicia acutifolia Helicia affinis Helicia albiflora Helicia amplifolia Helicia archboldiana Helicia arguta Helicia australasica Helicia blakei Helicia bullata Helicia calocoma Helicia cameronii Helicia dentellata Helicia elephanti Helicia excelsa Helicia falcata Helicia grayi Helicia insularis Helicia latifolia Helicia lewisensis Helicia loranthoides Helicia macrostachya Helicia maxwelliana Helicia microphylla Helicia neglecta Helicia nortoniana Helicia pallescens Helicia petiolaris Helicia platyphylla Helicia polyosmoides Helicia pterygota Helicia purpurascens Helicia recurva Helicia retusa Helicia rigidiflora Helicia sellae-montis Helicia sessilifolia Helicia sleumeri Helicia stelechantha Helicia stenophylla Helicia subcordata Helicia symplocoides Helicia wollastonii Helicia yangchunensis Helicia yingtzulinia Helicia moluccana Helicia rengetiensis Helicia vestita Helicia obovatifolia Helicia grandis Helicia pyrrhobotrya Helicia silvicola Helicia clivicola Helicia tsaii Helicia tibetensis Helicia hainanensis Helicia shweliensis Helicia longipetiolata Helicia formosana Helicia nilagirica Helicia reticulata Helicia cochinchinensis Helicia kwangtungensis Helicia dongxingensis Helicia woxvoldiana Helicia glabrescens Helicia retivenia Helicia carrii Helicia lauterbachiana Helicia fragilis Helicia graciliflora Helicia hypoglauca Helicia microcarpa Helicia microneura Helicia obtusata Helicia odorata Helicia olivacea Helicia oreadum Helicia biformis Helicia kjellbergii Helicia laiagamensis Helicia ledermannii Helicia cauliflora