Homalanthus A.Juss.

Genus

Angiosperms > Malpighiales > Euphorbiaceae

Characteristics

Monoecious or dioecious trees or shrubs with milky latex, commonly glabrous, rarely with a localized simple indumentum. Leaves alternate, long-petiolate, stipulate, peltate or not, simple, usually entire, penninerved, with a pair of glands at the apex of the petiole; stipules readily caducous. Inflorescences terminal or subterminal, pseudoracemose, solitary, usually bisexual, predominantly ♂ with 1 or more ♀ flowers at the base; ♂ bracts broad, with a pair of sessile glands at the base, 1–6-flowered, the lowest ones sometimes also each subtending a ♀ flower; ♀ bracts usually 1-flowered. Male flowers pedicellate; sepals 1 or 2, if 2 then imbricate and vertically disposed, giving the flowers a compressed appearance; petals 0; disc 0; stamens 5–50, free, filaments very short, anthers basifixed, bilobate, longitudinally dehiscent; pistillode 0. Female flowers long-pedicellate; calyx-lobes 2–3, open, not compressed; petals 0; disc 0; ovary 2(–3)-locular, with 1 ovule per locule; styles 2(–3), connate at the base, undivided or sometimes bilobate or shortly bifid, straight, the adaxial stigmatic surface running the length of the arms, canaliculate and minutely papillulose, the abaxial surface glandular or not. Fruit 2(–3)-lobed, subindehiscent or tardily loculicidally dehiscent; endocarp thinly crustaceous; columella small, subpersistent. Seeds ovoid-ellipsoid; endotesta crustaceous; caruncle fleshy, sometimes half enclosing the seed; cotyledons broad, flat.
More
Trees or shrubs, monoecious (sometimes one sex missing); latex white; indumentum absent [or consisting of multicellular, uniseriate, brownish hairs]. Apical buds enclosed by conspicuous large stipules. Leaves alternate; stipules caducous; petioles long, 2-glandular at apex or eglandular; leaf blade sometimes peltate, margin entire [to serrate], pinnately veined. Inflorescence terminal, elongate racemelike thyrses, unbranched. Flowers unisexual, apetalous, without disk. Male flowers 1-3(to many) per bract, pedicellate; calyx 1-or 2-lobed, free, bilaterally compressed; stamens 4-50; filaments nearly as long as anthers, free; anthers basifixed, extrorse, 2-thecate, longitudinally dehiscent; pistillode absent; receptacle flat to elevated. Female flowers usually 1 per bract, pedicellate; calyx 1-3-lobed, free, eglandular or glandular; ovary smooth, papillate to puberulent, with 2(or 3) locules; ovules 1 per locule; style short to distinct; stigmas 2(or 3), glandular below, apex often bifid. Capsule subglobose, regularly or irregularly dehiscent. Seeds 2(or 3) per fruit, reddish to brownish, flattened, partly to completely covered by a whitish arillode; testa crustaceous; cotyledon broad and flattened.
Plant monoec.; fls very small, in terminal racemes, apetalous. ♂ ∞ in upper part of raceme; sepals 1-2, minute; stamens us. ∞. ♀ few or solitary, at base; calyx 2-3-partite; ovary 2-3-celled, ovules 1 per cell; styles 2-3. Capsule fleshy, ind. or splitting into 2-3, 2-valved cocci. Aril fleshy. Shrubs or small trees with alt., entire, stipulate lvs. Some 20 spp. of Polynesia, Malaya, Australia. The N.Z. sp. endemic.
Monoecious shrubs or small trees; sap milky. Lvs stipulate, alternate, entire. ♂ and ♀ fls in the same or separate terminal racemes or spikes. ♂ fls: perianth reduced, of 1-2 segments; petals 0; anthers few to numerous; filaments free. ♀ fls: perianth reduced, in 2-3 segments; petals 0; ovary 2-3-celled, with 1 ovule in each cell; styles 2-3, undivided. Fr. fleshy, indehiscent or dehiscent.
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Hardiness (USDA) 9-12

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